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Order Dermaptera – Earwigs Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Dermaptera (Earwigs) Explanation of Names Dermaptera = "skin wings" Refers to the leathery texture of the forewings. Earwig is from Old English eár-wicga="ear-insect". Partridge (Origins--A Short Etymological Dictionary of Modern English, 1958) gives the meaning of Old English [i]wicga as beetle or worm, and mentions that folklore has it that the insect creeps into the human ear. Numbers Over 1000 earwig species worldwide, with about 20 species in North America. Nomina Nearctica lists 4 families: Anisolabididae (=Carcinophoridae) Forficulidae Labiduridae Labiidae More recent sources include 2 more families that Nearctica missed: Chelisochidae Pygidicranidae Size 6-35 mm long Identification Earwigs have slender, flattened bodies with bead-like antennae, and are easily recognized by the pair of large pincers (cerci) at the tip of their abdomens. Adult males have 10 abdominal segments (the pincers are one segment), while the females have 8. Habitat Earwigs tend to hide under debris during the day, but feed on plants, organic matter, and smaller insects at night. Food Plants, organic matter, other small insects. Life Cycle Simple metamorphosis with visible changes including increasing antennal segments and progressive wing development until sexual maturity. The mother cares for the eggs and nymphs. Simple metamorphosis progresses from the egg >>> nymph >>> adult. Remarks Earwigs were thought to crawl into people's ears at night, however this is an unfounded superstition. Earwigs are considered harmless to people, though they may emit a foul smelling liquid when disturbed. Larger earwigs might use their abdominal "pincers" to "bite" someone, but no venom is present. Though potentially unpleasant and perhaps disconcerting, no harm is done. Because earwigs hide during the day in dark places, they often get transported along with the object they hide in, and are easily overlooked. Most of our species are introduced, and new ones are constantly being introduced. See Also Rove beetles (Staphylinidae) - have no "pincers" at the tip of their abdomens. Print References Milne, pp. 407-410(1) Internet References Tree of Life Insects of Cedar Creek Dermaptera The Order Demaptera (earwigs) in Florida and the United States. List of Florida species, with pictures and a key to families. (pdf format) Wikipedia--Earwig--discusses etymology Earwigs of California (Order Dermaptera) 1975 California Insect Survey publication with species descriptions and a key for all California species. (pdf format) Works Cited Contributed by Richard Leung on 14 March, 2004 - 3:36pm Additional contributions by jvandyk, cotinis, Beatriz Moisset, Chris Wirth, Chuck Entz Last updated 8 February, 2009 - 11:08am Order Hymenoptera - Ants, Bees, Wasps and Sawflies Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Hymenoptera (Ants, Bees, Wasps and Sawflies) Synonyms and other taxonomic changes To see the taxonomic structure used on BugGuide.net for the Hymenoptera click here Explanation of Names From Greek, hymen, membrane, plus pteron (a), wing. Numbers Insects of Cedar Creek states there are over 70 families and about 18,000 species in North America. Arnett (1) gives a figure of 17,777 species, 2034 genera for North America. Size Most are of moderate size, but they range in size from miniscule (.2 millimeter) to 100 mm. Identification One of the most diverse groups of insects in form and life-history. Characteristics of Hymenoptera (1), (2): Typically two pairs of wings, with forewings usually larger than hindwings, but some groups (such as ants) wingless in most life stages Wings have few cross-veins, these are angled to form closed cells Antennae typically with 10 or more segments. Often 13 segments in male, 12 in female, but sometimes as few as 3 or up to 60 segments. Antennae longer than head, but usually not highly elongated (longer than head and thorax combined). Highly elongated in some parasitic groups. Females have prominent ovipositor, modified in some groups to be a "stinger", used to paralyze prey and in defense Chewing mouthparts, but some groups have a "tongue" used for lapping up fluids, such as nectar Complete metamorphosis Several groups highly social (eusocial), with separate reproductive and worker castes Range Worldwide Habitat Varied. Many adults are found on flowers. Food Food choice depends on particular genus and/or species. Many ants are predators or scavengers while others "milk" aphids and other insects for their sweet secretions, or cultivate fungus on cut leaves. Life Cycle All living ant species are eusocial (meaning, truly social). Complete Metamorphosis- egg>>>>>larva>>>>>pupa>>>>>adult See Also Velvet ants--Mutlillidae Print References Holldobler & Wilson (1) Milne & Milne, pp. 821-832 (2) Internet References AntWeb (California Academy of Sciences) AntData (Formicidae of the United States, David Lubertazzi, U. of Connecticut) Family Formicidae - various information, photos, classification tree (Animal Diversity Web, U. of Michigan) BioKIDS; Ants - information on ants, geared toward children (BioKIDS, U. of Michigan) Ants of Washington DC area - a list of 131 species (Georgetown U.) Myrmecos.net Zootaxa. Phylogeny and Systematics. Ants of Minnesota - Keys and Checklist (Carleton College) Works Cited Contributed by Richard Leung on 16 February, 2004 - 12:32pm Additional contributions by cotinis, Beatriz Moisset, Robin McLeod, Phillip Harpootlian, Chuck Entz Last updated 21 December, 2008 - 11:25am Order Orthoptera - Grasshoppers, Crickets, Katydids Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Orthoptera (Grasshoppers, Crickets, Katydids) Synonyms and other taxonomic changes Saltatoria Classification follows Orthoptera Species File; see Taxonomy Proposals topic here. Throughout the orthoptera, there is a tendency by some authorities to elevate some subfamilies to families, and by other authorities to demote families to subfamilies. Explanation of Names Orthoptera: from the Greek "orthos" (straight) + "pteron" (wing). Numbers Insects of Cedar Creek cites "1,082 species" in North America. Worldwide, more than 20,000 species. Identification Characteristics of Orthoptera include: hind legs long, modified for jumping forewings (tegmina) hardened, leathery, spread in flight, covering membranous hindwings at rest cerci (appendages at tip of abdomen) unsegmented pronotum usually with large descending lobes on sides hind coxae small and well-separated hind tibiae with two dorsal rows of teeth Two major taxonomic divisions: Caelifera - Grasshoppers and related families Ensifera - Long-horned Orthoptera, includes crickets and katydids Life Cycle Undergo Gradual Metamorphosis (paurometabolous development). Nymphs resemble small adults and typically develop external wing buds. They live in the same habitat as adults, typically eat the same food. egg>>>>nymph>>>>adult Range Varies; Worldwide Habitat Most species in grasslands, but some in forests, tundra, aquatic vegetation Season Spring-fall in temperate areas, some species present all year in southern regions Food Typically eat foliage of forbs, grasses. Some species take a variety of plants, while others are restricted to a few species of closely related plants. They often take dry plant matter from the ground as well, and most will scavenge weak or dead grasshoppers when plant food is scarce. Print References Capinera, Field Guide to Grasshoppers... (1) Internet References Tree of Life classification plus literature citations, synonym, and included taxa (Orthoptera Species File) Great Smoky Mountains National Park Orthoptera Checklist Insects of Cedar Creek Wikipedia: Orthoptera Works Cited Contributed by Troy Bartlett on 16 February, 2004 - 12:32pm Additional contributions by jvandyk, cotinis, John VanDyk, Robin McLeod, Jay Barnes, Wisconsin Oecanthinancy Last updated 17 February, 2009 - 10:56pm Order Lepidoptera - Butterflies and Moths Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Lepidoptera (Butterflies and Moths) Pronunciation Leh-pih-DOP-ter-ra Explanation of Names Lepidoptera means "scale wing", from Greek: lepis (λεπις / genitive form λεπιδος)- "scale" + ptera (πτερα)- "wing" Numbers Insects of Cedar Creek lists 75 families, and cites 11,293 species in North America. All-Leps lists 82 families and 12,423 species in North America. Identification Members of this order have four membranous wings (rarely wingless); hindwings are a little smaller than forewings, both largely or entirely covered with scales. Mouthparts adapted for sucking, the proboscis is usually in the form of a coiled tube. Moths usually have feathery antennae and most are active at night. They generally rest with their wings open, either flat or "tented" over the body. When they pupate above ground they generally form a protective cocoon around the pupa. This is made of silk, often combined with other natural materials such as leaves or their own body hair. The caterpillars of many species dig into the ground to pupate. Butterflies have thin antennae with "knobs" on the end and are generally active during the day. They rest with their wings closed above their bodies, and make a naked pupa also known as a chrysalis. Date/time-of-year and food plant (for caterpillars) are sometimes helpful in determining the species. Range Widespread Habitat Caterpillars are found in the habitat where their food is. Adults tend to be in the general area of their larval food plants, but many migrate over great distances. Season Caterpillars are active when their host plants are plentiful, which is often spring and summer. Adults are usually seen when there are flowers to feed on. Because they over-winter, though, there are individuals of some species to be found any time of the year, even in cold climates. Seasonal patterns are usually very specific for each species, and some can be told apart in the field only by when they're active. Food Most adult moths and butterflies feed on nectar sucked from flowers with their coiled, sucking mouthparts. In the process they may transfer pollen from one flower to another, and many plants depend on moths or butterflies for pollination. Most caterpillars eat the leaves of plants (usually very specific kinds for each species), but some carnivorous species have been documented. Life Cycle These insects undergo complete metamorphosis; that is, each individual goes through four stages: egg, larva (the caterpillar), pupa (chrysalis or cocoon), and adult. Egg>>>>larva>>>>pupa>>>>adult The larval stage does most of the eating and growing, with the adults often staying alive just long enough to mate and lay eggs. Remarks Contrary to popular belief, butterflies and moths will not die if the scales are rubbed off their wings. See Also Trichoptera (caddisflies) tend to have hairs rather than scales on their wings; no coiled proboscis Print References Borror and White, page 218 (1) Internet References Tree of Life Insects of Cedar Creek North Dakota State University's Key to the superfamilies of the Lepidoptera Information on butterflies and moths of North America North American Checklist with Synonymies (All-Leps) Links to Butterflies and Moths of Canada (CBIF) Zootaxa. Phylogeny and Systematics. Works Cited Contributed by Troy Bartlett on 16 February, 2004 - 12:32pm Additional contributions by jvandyk, cotinis, Hannah Nendick-Mason, John VanDyk, Beatriz Moisset, Robin McLeod, Lynette, Chris Wirth, Chuck Entz Last updated 14 February, 2009 - 2:33pm Order Diptera - Flies Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Diptera (Flies) Other Common Names The common names of true flies (order Diptera) are written as two words: crane fly, robber fly, bee fly, moth fly, fruit fly, etc. The common names of non-dipteran insects that have "fly" in their name are written as one word: butterfly, stonefly, dragonfly, scorpionfly, sawfly, caddisfly, whitefly, etc. Explanation of Names Diptera means "two wings" (Greek di two, plus pteron wing). The usage dates back to Aristotle, who noted, correctly, that they were different from typical insects with four wings and that no two-winged insect had a stinger (1). English fly originally signified any flying insect, likely the reason we have the more specific "fly" terms, such as butterfly noted above. Fly likely derives from Old English fleogan meaning to fly (Partridge, Origins--a Short Etymological Dictionary of Modern English). Numbers Arnett (2) notes 2,222 genera, 16,914 species in North America, with the number of described species increasing steadily (2) worldwide, 188 families with over 153,000+ described species Size 0.5-40 mm Identification Adult flies, except for wingless species, have two functional wings and two halteres. The halteres are club-like appendages that are essentially the modified hind wings. The only other adult insects that only have two wings in both sexes are the Strepsiptera, which have the front wings reduced rather than the hind wings. Males of some species of Mayflies and scale insects have only front wings. A few tiny parasitic wasps, e.g. Mymarommatidae, have their hind wings reduced, but these can be distinguished from flies as the wasps have only one vein in their front wings and flies always have two or more veins in their wings as long as their wings are membranous. Range Cosmopolitan Habitat Diverse. Adults can be collected in open landscapes, such meadows, at the edges of forests, inside forests, in the forest canopy, in stables or houses, or even in caves. Others can be found in coastal habitats or other places near water. For example, mosquitoes lay their eggs in water. Food Varies; some are parasitic like mosquitoes, whereas others feed on plant debris and decaying matter. Life Cycle Complete metamorphosis = egg>>>>larva>>>>pupa>>>>adult Print References Marshall, p. 591 (1) Arnett, order #29, pp. 835-920 (2) Internet References Tree of Life Insects of Cedar Creek Diptera.org Herschel Raney's Diptera keys A family key to the larvae of commonly collected flies found in aquatic habitats in Saskatchewan Anatomical Atlas of Flies (Need Broadband) Biodiversity Explorer Zootaxa. Phylogeny and Systematics. Paleontology of Diptera Order Coleoptera - Beetles Classification Kingdom Animalia (Animals) Phylum Arthropoda (Arthropods) Class Insecta (Insects) Order Coleoptera (Beetles) Other Common Names Grubs (larvae only) Pronunciation Coleoptera - Co`le*op"te*ra (coe-lee-OP-ter-rah) Synonyms and other taxonomic changes The Coleoptera have undergone A LOT of recent taxonomic changes, as reflected in American Beetles. Many groups that were once at family rank are now considered subfamilies and vice versa. If you can't find your group, search with a wildcard, e.g. Pselaphi* - Pselaphidae was commonly used but is now treated as the subfamily Pselaphinae in the Staphylinidae. Explanation of Names Coleoptera is derived from the Greek koleon (κολεον) meaning "sheath" and ptera (πτερα) meaning "wings" (though the word koleopteros (κολεοπτερος) was used to describe beetles as far back as Aristotle). This refers to the modified front wings (elytra) which serve as protective covers for the membranous hind wings. Eric Partridge, in Origins: a Short Etymological Dictionary of Modern English (1958), gives the meaning of English beetle as "the little biter", derived ultimately from Old English bitela, from the verb bitan (long -i) meaning, to bite. Numbers This is the largest order in the animal kingdom, with more than 350,000 described species worldwide, representing about 40 percent of known insects. Over 23,000 species in 131 families (1)(2) have been described in North America. Size Body length ranges from about 0.3 mm up to 200 mm (in tropical rhinoceros and goliath beetles) Identification Adult: elytra (forewings) horny or leathery, almost always meeting in a straight line down the back and covering the hindwing. The hindwings (the ones used in flight) are membranous, usually longer than the elytra, and folded beneath the elytra when not in use. Occasionally the elytra are short and do not cover the entire abdomen. 1 or both pairs of wings are rarely reduced or absent. The antennae usually have 11 segments, they rarely have more, often with 8–10, rarely with as little as 2; antennae variable in shape. The tarsi (the distal part of the legs) usually 3 to 5 segmented. Abdomen commonly has 5 segments visible, sometimes with up to 8. Mouthparts of adults and most larvae adapted for chewing. Larva: variable in form, hardness of body, and development of appendages, but commonly with hardened (sclerotized) head capsule, 3 pairs of thoracic legs, no wings, and soft body The order is divided into four suborders, with Polyphaga having the vast majority of species. Range cosmopolitan Habitat Members of this order can be found almost everywhere. Season All Food All sorts of plant and animal matter. Life Cycle Metamorphosis is complete; that is, the young go through four stages: egg, larva, pupa, and adult. Remarks Many species of plant-feeding beetles are serious pests of agricultural and forestry industries, and a number of predaceous beetles are considered beneficial to those same industries because they help to reduce prey populations. Some beetles feed on museum specimens, clothing, and stored food. A few species transmit bacterial and fungal diseases to plants. Print References Taxonomy is based mostly on American Beetles (1) (2) - see additions & corrections from the editors. See book references here. Internet References Families of Coleoptera same classification as used in American Beetles - with updates/exceptions as noted here Beetles of West Virginia, good on Appalachian species of Coleoptera Wikipedia article general oveview covering several topics (The Free Encyclopedia, wikipedia.org) Tree of Life overview of characteristics and classification with many references and links to related sites (David Maddison, U. of Arizona) Insects of Cedar Creek brief overview of classification plus links to adult images of many species (John Haarstad, U. of Minnesota) Museum Comparative Zoology (MCZ) type database - Harvard's searchable image database of type material Families of Beetles An Argentinian site containing photos representing every beetle family, along with technical descriptions (in Spanish) and subfamily lists