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POPULATION DYNAMICS Mrs. Cummings CP Biology POPULATION BASICS A population is a group of organisms of the same species living within a certain area. (Members of the same species are able to reproduce and have fertile offspring) The population size is the number of organisms within the group. The population density is the number of organisms per unit area. POPULATION GROWTH Population growth is the increase in the size of a population over time. As the organisms within a population reproduce, the population size increases. To graph population growth: X-axis = time Y-axis = population size POPULATION GROWTH POPULATION GROWTH Under ideal conditions, a population would grow rapidly and keep increasing toward infinity. Ideal conditions include favorable temperature, unlimited food/water, & lack of disease or predators. Biotic potential is the highest possible rate of reproduction for a population under ideal conditions. UNLIMITED POPULATION GROWTH: J-SHAPED CURVES Under ideal conditions, a population that starts out with just 1 or 2 individuals will increase in size slowly at first, then more quickly as more individuals become capable of reproducing. When graphed, this type of population growth resembles the letter J and is therefore called a J-shaped curve or exponential growth curve. UNLIMITED POPULATION GROWTH: J-SHAPED CURVES The size of the population is always increasing. The rate at which the population is growing is also increasing. You can tell how fast the population is growing by how steep the line is. LIMITING FACTORS In reality, most populations do not exhibit Jshaped growth curves. Limiting factors are circumstances that slow down population growth, preventing populations from reaching their biotic potential. Examples: Food, water, and space availability; predator species; unfavorable temperatures, etc . Limiting factors can be either density- DENSITY-DEPENDENT LIMITING FACTORS: S-SHAPED CURVES When the population size is small, densitydependent limiting factors have little effect, and the population can grow at a fast rate. As the population increases in size and density, density-dependent limiting factors begin to have more of an impact, and the growth rate of the population slows down. Results in S-shaped curve, also called a population growth curve or logistical growth curve. Density-dependent limiting factors include things like food, water, shelter, and predators. DENSITY-DEPENDENT LIMITING FACTORS: S-SHAPED GROWTH CURVES DENSITY-DEPENDENT LIMITING FACTORS: S-SHAPED CURVES The carrying capacity of a population is the largest sustainable size that it can reach in its given environment (such as a pond or woodland) under a particular set of conditions. The population can no longer grow because it has reached the maximum size that its environment can support. DENSITY-DEPENDENT LIMITING FACTORS: S-SHAPED CURVES At carrying capacity, the size of the population levels off. This is only possible when the birthrate equals the death rate of the population. DENSITY-INDEPENDENT LIMITING FACTORS & THEIR GROWTH CURVES Density-independent limiting factors will inhibit the growth of a population whether it is large or small. When a density-independent limiting factor affects a population, the population size may drop suddenly. The growth curve resembles a J-shaped or Sshaped curve until the drop-off occurs. These populations never reach their carrying capacity. DENSITY-INDEPENDENT LIMITING FACTORS & THEIR GROWTH CURVES The most common densityindependent limiting factors are seasonal changes in weather, especially temperature. POPULATION GROWTH RATE CURVES Growth rate is a measure of how fast a population is growing. Recall that in a population growth curve, the growth rate is indicated by the slope (steepness) of the line. A population growth rate curve shows how the rate of population growth changes over time. In a population growth rate curve, time goes on the x-axis, and growth rate goes on the y-axis. POPULATION GROWTH RATE CURVES Growth Rate Curve for a Population of Yeast Growth Rate (cells/hour) 200 180 160 140 120 100 80 60 40 20 0 1 2 3 4 5 6 7 8 9 10 11 Time (hours) 12 13 14 15 16 17 18 19 POPULATION INTERACTIONS The limiting factors that control population sizes often involve interactions among organisms. These include interactions between members of the same species and interactions between different species. PREDATOR-PREY RELATIONSHIPS Predation is the feeding of one organism on another. By eating prey, predators act as a limiting factor on prey populations. Predation is a density-dependent limiting factor. The greater the density of the prey population, the greater the likelihood of a predator finding one and eating it. But, since they are a food source for predators, prey species are also a limiting factor on predator populations. Also a density-dependent limiting factor! PREDATOR-PREY RELATIONSHIPS TIME PREDATOR-PREY RELATIONSHIPS Predation can prevent overpopulation of prey species. Predators help keep prey populations at sizes that the environment can sustain (carrying capacity!!) PARASITE-HOST RELATIONSHIPS Parasitism is when a small organism lives on or inside of a larger organism and causes it harm. The smaller organism is the parasite, and the larger organism is called the host. Parasitism can be a limiting factor on host populations. It is density-dependent because the parasite is more likely to be passed from one organism to another in a dense host population than in a more spread-out host population. SYMBIOTIC RELATIONSHIPS Parasitism is one type of symbiotic relationship. A symbiotic relationship or symbiosis is a close interaction between organisms of two different species. There are three types of symbioses: Parasitism- One organism benefits, the other is harmed Commensalism- One organism benefits, the other is not affected Mutualism- Both organisms benefit SYMBIOTIC RELATIONSHIPS Parasitism – One organism (the parasite) benefits while the other (the host) is harmed. Even though the parasite harms the host, killing the host is not its goal. The host is its home; if the host dies, it will need to find another host, which is risky! Caterpillar & wasp larvae Human tapeworm SYMBIOTIC RELATIONSHIPS Commensalism – One organism benefits; the other is neither harmed nor helped. Barnacles and sea turtle Sucker fish and shark SYMBIOTIC RELATIONSHIPS Mutualism – Both organisms benefit. Flower and butterfly Bird and crocodile DISEASE AND POPULATIONS Disease is a density-dependent limiting factor. The denser the population, the greater the chance that disease will spread. Humans have used diseases to purposely control populations of certain species. DISEASE AND POPULATIONS Studying how diseases spread through populations (epidemiology) can be complicated! For example, the bubonic plague spread rapidly through Europe in the 14th century and by some estimates killed over half the population! The plague is caused by a bacterium, which lives on fleas, which are carried by rodents & other animals… So the spread of the plague through human populations also depends on the population densities of fleas, rats, and other animals. COMPETITION FOR RESOURCES Usually many different species live in the same area at the same time. Often several species living in a given area share the same food source. For example, mice are food for owls & other birds, snakes, foxes, and many other animals. Interspecific competition is competition between members of different species in an area. Intraspecific competition is competition between members of the same species in an area. INTERSPECIFIC COMPETITION Competition between populations of different species. Interspecific competition is a density-dependent limiting factor. It may result in one of three possible outcomes: Extinction of one species in the area Movement of one population to another area Adaptation of one species due to natural selection (this occurs over many generations!) INTERSPECIFIC COMPETITION: EXTINCTION When 2 or more species living in the same environment compete for the same resources, one possibility is that one of the species will die out and become extinct in that environment. Paramecia (singular: paramecium) are single-celled organisms that live in large numbers in aquatic environments. In a classic lab experiment, two different species of paramecia are placed in a culture dish together with a limited supply of food. The results are shown on the following slide: INTERSPECIFIC COMPETITION: EXTINCTION When grown separately, populations of P. aurelia and P. caudatum both live. Growth of P. aurelia When grown together, P. caudatum dies out. P. aurelia & P. caudatum grown together Growth of P. caudatum 300 600 600 500 Number per 0.5 mL 500 Number per 0.5 mL Number per 0.5 mL 250 400 300 200 400 300 200 200 150 100 50 100 100 0 0 0 0 0 2 4 6 8 10 12 14 16 0 2 4 6 8 10 12 14 16 Day Day 2 4 6 8 10 12 14 16 Day INTERSPECIFIC COMPETITION: EXTINCTION Interspecific competition between P. aurelia and P. caudatum results in extinction of the P. caudatum P. aurelia & population. Growth of P. aurelia P. caudatum grown together Growth of P. caudatum 300 600 600 500 Number per 0.5 mL 500 Number per 0.5 mL Number per 0.5 mL 250 400 300 200 400 300 200 200 150 100 50 100 100 0 0 0 0 0 2 4 6 8 10 12 14 16 0 2 4 6 8 10 12 14 16 Day Day 2 4 6 8 10 12 14 16 Day INTERSPECIFIC COMPETITION: MOVEMENT A second possibility is that one species will move into another, usually nearby environment. • For example, several different species of Maine warblers make their nests in the same kind of spruce tree, but each species lives only within a certain zone of the tree (such as the crown, lower branches, or outer branches). INTERSPECIFIC COMPETITION: ADAPTATION The third possible outcome of interspecific competition is that rapid evolution will occur in one of the species, causing it to acquire different traits so it no longer has to compete with the other species. The finches that Darwin studied had developed different-shaped beaks suited for different types of seeds. Even though they live in the same area, they do not compete for the same food source because of these adaptations. This occurs because of random mutations and takes many generations! INTRASPECIFIC COMPETITION Competition between members of the same species. Intraspecific competition is a density-dependent limiting factor. Species have several ways of avoiding competition within their own populations: • • • Life Cycles & Life Spans Dominance Role Separation • • • Behavioral & Physiological Changes Emigration Territoriality LIFE CYCLES & LIFE SPANS The life cycle of a species can help reduce competition between members of that species. Frogs and tadpoles do not compete because their food and habitats are different. In many plants and fungi, dispersal of seeds or spores to distant locations helps prevent competition among members of the same species. Life spans may also help avoid intraspecific competition. In many insect species, adults die shortly after the young are produced, so old and young do not have to compete with each other. DOMINANCE A social hierarchy or pecking order is a chain of command that develops within a population and helps prevent conflict between members of the same species. The term “pecking order” comes from chickens pecking at each other to establish dominance. Male hierarchies also form in some populations, in which males establish dominance by fighting, especially with horns or antlers. Often the dominant male is the only one who mates, which helps prevent overbreeding. ROLE SEPARATION Competition is reduced in societies in which each member has a definite role. Occurs in many insect populations such as bees, ants, and termites BEHAVIORAL & PHYSIOLOGICAL CHANGES Overcrowding can cause changes in both behavior and physiology in members of a population. For example, overcrowded rats become increasingly aggressive to the point that more deaths occur than normal (behavior). Stress from overcrowding can also cause hormonal changes that reduce litter size (physiology). As a result of these changes, birthrate decreases while death rate increases. This reduces the population size to a number the environment can support. EMIGRATION Emigration is when some members of an overcrowded population relocate to a new area. Some members remain in the original environment. Results in 2 low-density populations. Bee example from Chapter 27 Intro TERRITORIALITY Territoriality is when different members of the same species occupy and defend different territories. By staying separated, competition for food and other resources is avoided. Males that control a territory are often the only ones to mate, which helps limit population growth. THE HUMAN POPULATION Human population growth resembles a J-shaped curve. THE HUMAN POPULATION THE HUMAN POPULATION Growth rate is declining. The growth rate is much higher in developing countries than in industrialized countries. Population size is still increasing. THE HUMAN POPULATION AGE STRUCTURE DIAGRAMS (POPULATION PYRAMIDS) To predict population growth, it is useful to know the age of everyone in the population. People are often categorized by age groups. An age structure diagram (also called a population pyramid) is a tool used to visually display how many people within a given population are in each age group. The proportion of females of reproductive age is especially important. AGE STRUCTURE DIAGRAMS (POPULATION PYRAMIDS)