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Classification of Organisms 1st system of Classification was designed by Aristotle, all organisms were categorized as walks on land, swims in water, or fly in the sky. Carolus Linneaus’ system replaced Aristotle’s using a series of categories. Common features were used to place them in categories. The more categories organisms share the more closely related they are. Binomial Nomenclature- system of assigning 2 names to every species Scientific Name- genus and species, ex Homo sapien (human), Canis lupus (wolf) Traditional Classification system: What Linnaeus also did was systematically categorize all known organisms. Linnaeus came up with a hierachy of ways to classify plants and animals. The different levels are called taxa (plural of taxon). The different taxa are: Kingdom Phylum Class Order Family Genus -part of the scientific name Species -part of the scientific name Domain and Kingdom are the broadest of the taxa, all animals are in Kingdom Animalia, all plants are in the Kingdom Plantae. The taxa Domain is the newest addition, and was created to differentiate the two very different groups of bacteria. Phyla are slightly less broad. There are usually a few phyla in each kingdom. Species is the most restricted. Taxonomist classify new organisms. Historically classification was based on easily observable structures, and gradually this was extended to microscopic and electronmicroscopic detail. The recent advances in embryology and molecular biology have given new tools such as patterns of life cycle, larval development, and gene sequences. These have often led to radically different phylogenies (e.g. humans should really be the "third chimpanzee"). After they assign each new organism a scientific name, taxonomist work closely with Evolutionary Biologist to develop the organism’s phylogeny (evolutionary history this is part of the science of systematics, the study of biodiversity and its classification. Using comparative anatomy, embryology, fossil record, and molecular biology, scientist can search for common ancestors and develop a cladistic analysis or phylogenetic tree. The cladogram depicts patterns of shared characteristics among taxa and forms the basis of a phylogentic tree. A clade, within a tree, is defined as a group of species that includes an ancestral species and all its descendants. 1 Molecular Clocks help track evolutionary time, this method is used to measure the absolute time of evolutionary change based on the observation that some genes and other regions of the genome appear to evolve at constant rates. The rate of evolution of DNA sequences varies from one part of the genome to another. Comparing these sequences and analyzing the number of differences given scientist and idea of when they had a common ancestor and when they diverged. DNA that codes for ribosomal RNA changes relatively slowly and is used to investigate relationships in taxa that diverged hundreds of millions of years ago. DNA that codes for mitochondrial DNA evolves rapidly and is used to investigate more recent evolutionary events, for instance the evolution and migration of man. 2 Helpful resource: http://evolution.berkeley.edu/evolibrary/article/phylogenetics_01 Dichotomous key, is a tool used to identify an unknown species. It is written using two questions at a time with a directive. The end result is the user can give the scientific name of the species after looking at specific characteristics and narrowing down the choices. The Evolution of Complexity: single cell prokaryote to multicelluar eukaryotes Earth is approximately 4.5 billion years young The first living things, prokaryotic bacteria cells, are found in 3.5 billion year old strata/ fossil record The oldest fossils of Eukaryotic cells appear in 2.1 billion year old strata Between 635- 530 million years ago the fossil record shows the diversity of algae and small Biogenesis: theory that life comes from life How did life begin? The hypothesis of Oparin and Haldane were tested by Stanley Miller in 1953, Miller was able to recreate anaerobic conditions similar to those hypothesized by Oparin and synthesize organic molecules from inorganic using an apparatus he built. Research on prebiotic origin of organic compounds continues. Virus: Not a living cell, but rather a protein coat surrounding genetic material either DNA or RNA.(retroviruses contain RNA and reverse transcriptase to transcribe RNA into DNA ex. HIV) Two types of viruses: 1. reproduce using Lytic cycle, the virus DNA takes over the host cell to produce copies of the virus and ends in the lysis or death of the host cell. 2. Lysogenic cycle, the DNA is incorporated into the genome of the cell.. Modern Classification system: Domain, Kingdom, Phylum, Class, Order, Family, Genus, species Domain was added when it was discovered that bacteria (Monera) were not that closely related to each other, in fact some bacteria have more in common with eukaryotes than other bacteria. 3 Domains: Archea, Bacteria, Eukarya 6 Kingdoms: Archeabacteria, Eubacteria, Protista, Fungi, Plantae, Animalia 3 Common Characteristics of each Kingdom: Kingdom Archeabacteria Eubacteria Protista Fungi Plant Animal Characteristics Prokaryotes, extreme environments: deep ocean, volcanic ridges, geysers, hot springs, salt lakes, heterotrophic or autochemotrophs, no peptidoglycan in cell membrane, has histones and introns associated with DNA/chromosome Prokaryotes, 3 common shapes rod, sphere, spiral, may be autotrophic or heterotrophic, has peptidoglycan which is how they are classified (Gram + or Gram -) no histones or introns associated with their DNA/chromosome ex. E.coli, Streptococcus Eukaryotic cells (with nucleus) typically single celled but may form colonies, Autotrophic- Algae, Heterotrophic Protozoa and fungus-like Eukaryotic multicelled, Heterotrophs, cell wall of chitin, reproduce with spores, can not move Eukaryotic multicelled, Autotrophs, cell wall of cellulose, reporoduce with spores or seeds, can not move Eukaryotic multicelled, Heterotrophs, can move, no cell wall, can be invertebrate or vertebrates Domain: the broadest taxa Characteristic Nucleus Membrane bound organelles Introns Histone proteins associated with DNA Circular chromosome Peptidoglycan in cell wall Hydrocarbon content Bacteria No No No No Archaea No No Yes Yes Eukarya yes yes yes yes Yes yes Un-branched hydrocarbons Yes no branched hydrocarbons no no Un-branched hydrocarbons 6 Kingdoms: 1. Archeabacteria- prokaryotes, most ancient and extreme, living in the harshest environments, include methanogens (anaerobic/no oxygen), thermoacidophiles (hot), and extreme halophiles (salty). No Peptidoglycan in cell wall 2. Eubacteria- prokaryotes, true bacteria, have peptidoglycan, classified by their shape and response to gram staining (gram positive high level of peptidoglycan, gram negative less peptidoglycan) 4 Eubacteria Shapes: Sphere- Cocci, can occur in chains Streptococcus pneumoniae which can cause Strep throat or Scarlet Fever, or grapelike clusters Staphylococcus aureus causes Toxic Shock Syndrome Rod- Bacillus ex. Escherichia coli (E.Coli), Lactobacilli causes tooth decay, botulism, typhoid fever, anthrax Spiral- Spirilla ex Vibro (curved) ex Cholera, Spirillum (thick spiral), Spirochete (thin spiral) ex Treponema pallidum causes Syphilis and Lyme disease Cyanobacteria- photosynthetic bacteria, contain pigments- yellow, red, brown, green, and even black, some are blue-green. Like plants they thrive on Phosphates and Nitrates and can have a Population Bloom if water ways become loaded with fertilizer (process called Eutrophication) Nitrogen-Fixing Bacteria- grow symbiotically in the roots of plants, fixing atmospheric nitrogen into ammonia so plants can utilize it. Asexually reproducing – bacteria still have a tremendous amount of diversity. They have Rapid reproduction, rapid mutation rates, and genetic recombination due to the following: Transformation: taking up DNA fragments from the environment Conjugation: exchanging DNA plasmids with another bacteria via a cytoplasmic bridge Transduction: a virus delivers genetic material to the bacteria, which gets incorporated into the genome of the bacteria All Prokayotes: Reproduce asexually By binary fission Single Celled organisms Cell Wall Single Strand of DNA Some: Autotrophic or Heterotrophic some produce toxins Some movement using flagella, slime, spiral motions Some produce Endospores and go dormant during extreme/hostile conditions Bacteria can be helpful: fermentation process, digestion, biotechnology like bioremediation where bacteria help digest petroleum products or heavy metals in water or soil, nitrogen-fixing a symbiotic relationship some bacteria have with plants, photosynthetic bacteria fill water ways and oceans and act as CO2 sponges and release oxygen also. Bacteria can be killed using antibiotics which destroy the cell wall, Gram Negative bacteria have and extra lipid layer that stops antibiotics from entering the bacteria Eukaryotes evolved about 1.2 billion years ago. Endosymbiosis: theory that bacteria gave rise to eukaryotic cells by engulfing bacteria capable of photosynthesis and cellular respiration. This is the origin of the organelles mitochondria and chloroplast 5 3. Protistia Kingdom of Mostly single celled organisms Categorized by their likeness to 3 other kingdoms All Eukaryotic some Autotrophic and some Heterotrophic 3 classifications: Algae or Plant-like, Autotrophic, classified by pigment, cell wall, can join to form elaborate colonies and multi-cellular structures (kelp and seaweed) gave rise to land plants. Chrysophyta- Golden Algae ex diatoms Pyrrophyta- bioluminescent ex. Dinoflagellates - cause red tide Euglenaphyta- ex Euglena Rhodophyta- red Phaeophyta- brown, seaweed and kelp Chlorophyta- green, ex Volvox, Spirogyra Protozoa or Animal-like, classified by mode of movement Sarcodina ex. Amoeba- move by pseudopodia “false foot” Ciliophora ex Paramecium- move by cilia, tiny hairs Zoomastigina ex Trypanosoma (African Sleeping Sickness) moves by flagella Sporozoa, are the parasitic animal like protista, include Plasmodium which causes Malaria Fungus-like are all Heterotrophic with no cell wall, absorbing nutrients directly through cell membrane Include Slime Molds and Downy Mildews Multicelled Eukaryotes-The colonization of land occurred about 500 million years ago when plants, fungi, and animals began to appear on Earth 4. Fungi All Eukaryotic, multicellular, heterotrophs (saprotrophs or detrivores living off dead and decaying organisms or decomposers), cell wall made of chitin (type of protein), and classified by reproductive structures. Mycoses is the term for Fungal Infection, Fungus destroys the cells around the infection site, to fight fungal infection you have to destroy the cell wall Fungi are important part of an ecosystem because they recycle nutrients 6 Club Fungi – Basidiomycota , Basidia, ex Mushrooms Sac Fungi- Ascomycota, Asci, ex. Yeast, and Athletes Foot Fungus Zygote Fungi – Zygomycota, Sporangia, ex. Bread molds Imperfect- Deuteromycota, reproduce asexually, ex Penicillian Lichen- Mycophycophyta, symbiotic relationship between fungus and photosynthetic cells of cyanobacteria or algae. 5. Plantae Multicellular, Eukaryotic, Cell wall made of cellulose, Autotrophs, not capable of movement, reproduce sexually some produce spores and others seeds. Most have vascular tissues which include roots, stems, and leaves, and specialized tissues called xylem and phloem. 9 divisions of Plant Kingdom: Spore producers Bryophyta only nonvascular plants, ex moss and liverwort (gametophyte dominant stage of life cycle know as alternation of generations) Lycopodophyta ex. 1,000 species of Club and Spike moss Sphenophyta ex. 10 species of Horsetails Filicinophyta ex 11,000 species of Ferns (sporophyte dominant stage) Seed producers Gymnosperms: reproductive structure is the cone , “naked seed” o Cycadophyta 200 species of these tropical plants o Ginkgophyta 1 species remains the Gingko tree o Coniferophyta 600 species of conifers, pine, spruce, cypress, juniper, fir o Gnetophyta 70 species of these arid/semi-desert dwelling plants Angiosperms: reproductive structure is the flower, seeds are surrounded by fleshy or dry fruit that ripen to encourage animals to disperse the seeds. 30,000 or more identified species maple, hickory, oak, aloe, roses, tulips, dogwood, magnolia, corn, beans, tobacco, apple tree, pecan tree Monocots – corn, with one seed leaf or cotyledon and parallel veins on the leaf, Dicots- beans, with two seed leafs and branched veins on the leaf Symbiotic relations ships: Plants and Bacteria Flower patterns and scent attract pollinators Fruits promote seed dispersal by offering a sweet reward to animals. Plant Adaptations include hormones that control, phototropism, geotropism, and thigmotropism, specialized leaves (needles, spines, fuzzy, waxy), specialized seed capsules (wind, float, food, wings, burrs, sticky) 7 6. Animalia multicellular, eukaryotic, no cell wall, heterotrophs, capable of movement, reproduce sexually, body symmetry (radial or bilateral) Invertebrates: 8 phyla Porifera- two cell layers, collar cells w/flagella, filter feeders, ex. sponges Cnidarians- Stinging cells, radial symmetry, simple nervous system only stimulus and response, central cavity only one body opening, ex. jellyfish, coral, hydra, sea anemone Plathyhelmenthes, flat worms, only one body opening, three cell layers, but no true body cavity (acoelomates) ex. Tapeworm, Fluke, Planarian, Marine worms Nematode, round worms, first with two body openings, 3 cell layers, (pseudocoelomates) many parasitic, Heartworm, Roundworm, Vinegar Eel Annelid, segmented worms, two body openings, 3 cell layers, (coelomates true body cavity) beginning of circulatory system, and digestive system (crop), ex. Earthworm and Leech Mollusk, soft body, more complex body systems developing, eyes and nervous system, levels of communication beyond stimulus and response, 3 classes include Gastropod (snails and slugs), Cephalopod (squid octopus cuttlefish and chambered nautilus), Bivalve (oyster, clam, mussel, scallop) Arthropod, jointed legs, exoskeleton, body segments (head, thorax, and abdomen), metamorphosis (complete or incomplete) 4 classes include Insect 6 legs (ant, grasshopper, beetle, bee, wasp) Arachnid 8 legs (spider, tick, horseshoe crab), Crustacean 10 legs (shrimp, lobster, barnacles, crayfish), and Myrapods many legs, centipede and millipedes Echinoderm, spiny skin that covers and endoskeleton, complex regeneration capacity, found only in marine environments, radial symmetry, ex Sea urchin, Sand dollar, Starfish, Sea cucumber o Echinoderms and Chordates share a developmental characteristic: they are deuterosomes which means during the gastrula stage and the formation of the coelom or body cavity the anus is formed first, the other bilateral inverts are protostomes and they develop their mouth first. Chordata, notachord, complex body systems, sexual reproduction (internal or external fertilization) Two invertebrate subphyla- tunicates and lancelets separated from the other Chordata 550 million years ago during Precambrian 8 Subphyla vertebrates with 5 classes: o Fish, moist skin covered in scales, gills to breathe, 2 chambered heart, cold blooded, 3 types: 1.jawless, ex. Lamprey 2. Bony (Osteichthyes) ex Trout, Salmon, Bass, Catfish, Grouper, Tarpon. and 3. Cartilaginous (Chondrichthyes) ex. Sharks and Rays o Amphibian, cold blooded, 3 chambered heart, born in water, develop lungs, smooth moist skin, ex. salamander, newts, skinks, frogs, and toads (only dry skin) o Reptile, dry leathery skin with scales, cold blooded, 3 chambered heart except for crocodile and alligator which have 4, amniote/terrestrial eggs, ex. Lizards, snakes, turtles, alligator, crocodile o Bird, (Aves) warm blooded, 4 chambered heart, hollow bones, body with feathers o Mammalia, warm blooded, 4 chambered heart, milk producers, body with hair/fur o Placental (live birth), Marsupial (pouch) ex. Kangaroo, and Monotremes (egg) ex Platypus o Humans belong to the order Primates along with monkeys and gorillas. Common characteristics include: opposable thumbs, large brains and short jaws, forward looking eyes, flat nails, well-developed parental care, and complex social behavior. o Human alone have evolved increades brain volume, shortening of the jaw, bipedal posture, reduced size difference between sexes, and certain important changes in family structure. Animal Behaviors and Adaptations: Innate Behaviors and Learned Behaviors Innate: 1. Instincts- complex pattern of innate behaviors, reflexes, fight or flight, courtships, species recognition (language, song, flashes of pattern/light) 2. Territory- physical space needed for breeding, feeding, and shelter, organisms can expend a lot of energy defending territory some will fight to the death. 3. Migration- instinctive seasonal movement, response to a changing environment, includes hibernation (cold) and estivation (dry and hot) Learned: 1. Habituation- animal repeats a successful behavior, and does not repeat an unsuccessful behavior, birds learn which moths are poisonous by color and avoid eating them after becoming ill or getting a bad taste. Deer return to the same grazing field when successful. 2. Imprinting – salmon and turtles return to same stream or beach to lay eggs in which they hatched, the environment left an imprint or memory 9 Adaptations for Defense: 1. Mechanical- physical structures 2. Chemical- stinging sensations, poisons, bad taste, paralysis 3. Camouflage- color or pattern that blend into environment o Disruptive- ex zebra o Cryptic- ex chameleon and squid o Countershading- ex Fish have light belly and dark back 10