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CARDIORESPIRATORY ADAPTATIONS TO TRAINING During a single bout of exercise, the human machine is quite adept at adjusting its cardiovascular and respiratory functioning to adequately meet the heightened demands of active muscles. When these systems are faced with these demands repeatedly, such as on a daily basis when training, they adapt in ways that allow the body to improve its performance of endurance activity. For example, the miler can run a faster mile. The physiological and metabolic processes that bring oxygen into the body, distribute it, and allow it to be used by active tissues become and remain highly efficient at these tasks. In this chapter, we will examine adaptations in cardiorespiratory function in response to training, and how such adaptations affect an athlete’s endurance capacity and performance. Cardiorespiratory endurance is probably the least understood component of a total training program. The training programs for many nonendurance athletes completely ignore the endurance factor. This is understandable, because for maximum improvement in performance, training should be higbly specific to the particular sport or activity in which the athlete participates, and endurance is frequently not recognized as important to nonendurance activities. The reasoning then is "Why waste valuable training time if the result is not improved perfonnance?" The problem with this reasoning is that often, although it might not be obvious, a nonendurance sport does indeed have an endurance, or aerobic, component. For example, if you play football, you and your coach might fail to recognize the importance of cardiorespira~ tory endurance as part of your total training program. From all outward appearances, football is an anaerobic or burst-type activity, consisting of repeated bouts of high-intensity work of short duration. Seldom does a run exceed 40 to 60 yd (37 to 55 m), and even this is followed by a substantial rest interval. The need for endurance is not readily apparent. What you and your coach might fail to consider is that this burst-type activity must be repeated many times during the game. With a high endurance level, the quality of your burst activity could be maintained throughout the game, and you would still be relatively fresh during the fourth quarter. ' Sport scientists are beginning to realize the importance of endurance training for almost all types of sports or activities: . Burst-type, such as football,basketball and some parts in dance . Moderate-intensity and skilled, such as baseball ,golf and dance . Endurance-type, such as running. cycling and swimming Many authorities now believe football teams that fall apart in the final quarter have ignored the endurance component in their training programs, A similar case can be made for athletes in most sports. But before we look specifically at how endurance can improve performance, we must understand what endurance is. Endurance Endurance is a term that describes two separate but related concepts: muscular endurance and cardiorespiratory endurance. Each makes a unique contribution to athletic performance, so each differs in importance to different athletes. For sprinters, endurance is the quality that allows them to sustain a high speed over the full distance of, for example, the 100- or 200-m race. This quality is muscular endurance-젳the ability of a single muscle or muscle group to sustain high-intensity, repetitive, or static exercise. This type of endurance is also exemplified by the weight lifter, boxer, and wrestler. The exercise or activity can be rhythmical and repetitive in nature, such as the bench-press for the weight lifter and jabbing for the boxer. Or the activity can be more static, such as a sustained muscle action when a wrestler attempts to pin an opponent to the mat. In either case, the resulting fatigue is confined to a specific muscle group and the activity's duration is usually no more than I or 2 min. Your muscular endurance is highly related to your muscular strength and anaerobic development. Whereas muscular endurance refers to the ability of individual muscles, cardiorespiratory endurance relates to the body as a whole. Specifically, it refers to your body's ability to sustain prolonged, rhythmical exercise. This type of endurance is typified by the cyclist, distance runner, or endurance swimmer who can complete long distances at a fairly fast pace. Your cardiorespiratory endurance is highly related to the development of your cardiovascular and respiratory systems, and thus your aerobic development. KEYPOINT CARDIOVASCULAR ENDURANCE IS THE ABILITY OF THE BODY TO SUSTAIN PROLONGED EXERCISE EVALUATING AEROBIC POWER To study training effetcs on endurance, we need a means for evaluating an individual’s endurance capacity which we can easily use to mintor her or his improvement during the training program. VO2 MAX: AEROBIC POWER VO2 max is defined as the highest rate of oxygen consumption attainable during maximal or exhaustive exercise. If you increase your exercise intensity beyond the point at which you reach VO2 max, your oxygen consumption will either plateau or decrease slightly. Reaching the plateau means that you can’t deliver oxygen as quickly as is needed to meet your muscles’ demands You can continue exercising for a brief period after reaching VO2 max by calling on your aerobic resreves, but these also have a finite capacity. With endurance training , more oxygen can be delivered and consumed than in untrained state. Previously untrained subjects show average VO2 max increases of 20% or more after a 6 month training program. These improvements allow you to perform endurance activities at a higher work rate or faster pace, improving your performance potential. OXYGEN TRANSPORT SYSTEM Oxygen transportation and delivery are major functions shared by both your car diovascular and respiratory systems, All components of these two systems that are related to the transportation of oxygen are collectively referred to as the oxygen transport system. The functioning of the oxygen transport system is defined by the interaction of the arterial venous oxygen difference (a—vo2: diff). Cardiac output 琴volume x heart rate) tells us (stroke how much oxygen-carrying blood leaves the heart in 1 min. The arterial venous difference, which is the difference between the oxygen content of the arterial blood and the oxygen content of the venous blood, tells us how much oxygen is extracted by the tissues. The product of these values tells us the rate at which oxygen is being consumed by the body tissues. CARDIOVASCULAR ADAPTATIONS TO TRAINING Numerous cardiovascular adaptations occur in response to training: - Heart size Stroke volume Heart rate Cardiac output Blood flow Blood pressure Blood volume SYNTHESIZING 1. Cardiovascular endurance refers to your body’s ability to sustain prolonged, rhythmical exercise. It is highly related to your aerobic development 2. Most sport scientists regard VO2 max- the highest rate of oxygen consumption, obtainable during maximal or exhaustive exercise- to be the best indicator of cardiovascular endurance 3. Cardiac output tells how much blood leaves the heart each minute, whereas “a VO2 max diff.” Indicates how much oxygen is extracted from the blood by the tissues. The product of these tells us the rate of oxygen consumption: VO2max- SV x HR x a VO2 maxdiff 4. The left ventricle undergoes the most change in response to endurance training 1. The internal dimensions of the left ventricle increase, mostly in response to an increase in ventricular filling KEYPOINT A stronger heart and the availability of a greater blood volume appear to account for the increases in resting, submaximal, and maximal stroke volumes following an endurance training program. Studies in which the oxygen consumption of the heart has been directly monitored have shown that the heart rate, both at rest and during exercise, is a good index of how the heart is working. Because active muscle requires more oxygen than resting muscle, it is not surprising that the heart’s oxygen consumption, and thus the amount of work it performs, are directly related to the heart’s consrtaction rate. RESTING HEART RATE The heart at rest decreases markedly as a result of endurance training. If you are a sedentary individual with an initial resting heart rate of 80 beats /minute, your heart rate will decrease by approximately 1 beat/minute each week for the first few weeks of training. Highly conditioned endurance athletes often have resting heart rates lower than 40 beats/minute, and some have values lower than 30 beats/minute! Bradycardia is a clinical term indicating a heart rate of less than 60 beats/minute. In untrained individuals, bradycardia is usually the result of abnormal cardiac function or a diseased heart. Therefore, it is necessary to differentiate between training-induced bradycardia, which is a natural response to endurance training, and pathological bradycardia, which can be a serious cause for concern. HEART RATE RECOVERY During exercise your heart rate must increase to meet the demands of your active muscles. When the exercise bout is finished, your heart rate does not instantly return to its resting level. Instead, it remains elevated for a while, slowly returning to its resting rate. The time it takes for your heart rate to return to its resting rate is called the heart rate recovery period. RESISTANCE TRAINING AND HEART RATE The reductions in heart rate that have been reported in resistance training are much less than the typical reductions obtained in endurance training.These changes appear to be dependent on the following characteristics of the resistance training program: - training volume - training intensity - training duration - length of rest periods bewteen stes - amount of muscle mass used SYNTHESIZING 1. Resting heart rate decreases considerably as a result of endurance training. In a sedentary person the decrease is typically about 1 beat/minute per week during initial training. Highly trained endurance athletes often have rest rates of 40 beats/minute or less. 2. Heart rate during submaximal exercise also decreases, often by 20 to 40 beats/minute following 6 months of moderate training A person’s maximal heart’s rate decreases proportionally with the amount of traing completed. 3. Maximal heart either remains unchanged or decreases slightly with training. When a decrease occurs , it is probably to allow maximum stroke volume to maximize cardiac output. 4. The heart rate recovery period decreases wit瑨 increased endurance , making this value well suited to tracking an individual’s progress with training. 5. Resistance training can also lead to reduced heart rates, however these decreases are not as reliable or as large as those seen with endurance training. MAXIMUM BREATHING CAPACITY TEST To better understand the concept of optimizing the heart rate- stroke volume relationship ce the greatest ventilation vol to provide the greatest cardiac output. BLOOD FLOW We know that active muscles need much more oxygen and nutrients. To meet these needs, more blood must be brought into these muscles during exercise. As the muscles become better trained, the cardiovascular system adapts to increase blood flow to them. Three factors account for this enhanced blood supply following training: 1. Increased capillarisation of trained muscles 2. Greater opening of existing capillaries in trained muscle൳ 3. More effective blood redistribution. To permit increased blood flow, new capillaries develop in trained muscles. The allows the blood to more fully perfuse the tissues.This increase in capillaries is usually expressed as an increase in the number of capillaries per muscle fiber. The existing capillaries in trained muscles can open up more, which increases blood flow through the capillaries and into the muscles. Because endurance training also blood because more blood is present in the system to begin with, so shifting more into capillaries will not severely compromise venous return. Blood flow to the active muscles can also be increased by a more effective blood redistribution of the cardiac output. Blood flow is directed to the active musculature and shunted away from areas that don’t need high flow. Venous compliance can also be decreased with endurance training as a result of increased because more blood is present in the system to begin with, so shifting more into capillaries will not severely compromise venous return. Blood flow to the active muscles can also be increased by a more effective blood redistribution of the cardiac output. Blood flow is directed to the active musculature and shunted away from areas that don’t need high flow. Venous compliance can also be decreased with endurance training as a result of increased venous tone. That means that the veins are not as easily distended by the blood, so less blood pools in the venous system, thereby increasing the amount of arterial blood available for working muscles. BLOOD VOLUME Endurance increases blood volume. This effect is greater with more intense levels of training. This increased blood volume iactually results from an increase in blood plasma volume. This is thought to be primarily caused by two mechanisms. First, exercise increases the release of antidiuretic hormone( ADH) and aldosterone.These cause the kidneys to retain water which increases blood plasma.Second, exercise increases the amount of plasma,particularly albumin. The plasma proteins are the major basis for the blood’s osmotic pressure. The result is that more fluid is retained in the blood. Thus both mechanisms work together to increase the fluid portion of the blood- the blood plasma. RESPIRATORY ADAPTATIONS TO TRAINING No matter how efficient the cardiovascular system is at supplying adequate amounts of blood to tissues, endurance would be hindered if the respiratory system didn’t bring in enough oxygen to meet oxygen demands.Respiratory system functioning usually does not limit performance because ventilation can be increased to a greater extent than cardiovascular function. But, as with the cardiovascular system, the respiratory system undergoes specific adaptations to endurance training to maximize its efficiency. LACTATE THRESHOLD Endurance training increases lactate threshold . In other words,, after training you can perform at a higher rate of work and at a higher absolute rate of oxygen consumption without raising your blood lactate above resting levels.The lactate threshold occurs at a higher percentager of VO2 max after training. This increase in the lactate threshold appears to be due to several factors: - a greater ability to clear lactate produced in the muscle an increase in muscle enzymes The net result is less lactate production for the same work rate. Synthesizing 1. Lactate threshold increases with endurance training, which allows you to perform at higher rates of work and levels of oxygen consumption without increasing your blood lactate above resting levels. Maximal blood lactate levels can be increased slightly. 2. The respiratory exchnage ratio decreases at submaximal work rates, indicating a greater utilization of free fatty acids, but increases at maximal effort 3. Oxygen consumption can be increased slightly at rest and decreased slightly or decreased or unaltered during submaximal exercise 4. VO2 max increases substantially following training, but the amount of increase possible is limited in each individual.The major limiting factor appears to be oxygen delivery to the active muscles. FACTORS AFFECTING THE RESPONSE TO AEROBIC TRAINING We must always remember that we are talking about adaptations in individuals, so not everyone will respond in the same manner.several factors must be considered that can affect individual response to aerobic training: - Heredity Age Gender Even with identical programs, not everyone will respond the same. Genetics accounts for much of this variation in response. SPECIFICITY OF TRAINING Physiological adaptations in response to physical training are highly specific to the nature of the training activity. The program must be closely mathched to the indivual needs to maximize the physiological adaptations to training, thereby optimizing the athlete’s performance. CARDIOVASCULAR ENDURANCE AND PERFORMANCE Cardiorespiratory endurance is regarded by many people as the most important component of physical fitness. It is an athlete’s major defense against fatigue, even in the more sedentary sports or activities. For any athlete, regardless of the sport or activity, fatigue represents a major deterrent to optimal performance. Even minor fatigue can have a detrimental effect on the athlete’s total performance: - Muscular strength is decreased Reaction and movement times are prolonged Agility and neuromuscular coordination are reduced Whole-body movement is slowed Concentration and alertness are reduced The decline in concentration and alertness associated with fatigue is particularly important. The athlete can become careless and prone to injury. IN REVIEW 1. Although VO2 max has an upper limit, endurance performance can continue to improve for years with continued training 2. An individual’s genetic makeup predetermines a range for his or her VO2max, accounting for 25 to 50% of the variance in VO2 max values. Heredity also largely explains individual variations in response to identical training programs. 3 Age- related decreases in aerobic capacity might partly result from decreased activity 4 Highly conditioned female endurance athletes have VO2 max values only about 10% lower than those of highly conditioned male endurance athletes. 5 To maximize the cardiorespiratory gains from training, the training should be specific to the type of activity that the athlete performs. Resistance training in combination with endurance training does not appear to restrict improvement in aerobic capacity and may increase short-term endurance