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Phylum Cnidaria • Notice the trend toward complexity as we go through each phylum! • Ny-dare-ee-ah • The phylum Cnidaria is comprised of the following classes: Hydrozoa (hydras, obelia, P.M.O. War), Scyphozoa (jellyfish) and Anthozoa (anemones and corals). • Comes form the Greek knide for nettle • Cnidarians have radial symmetry • The major advancement is the development of true tissues. Phylum Cnidaria • Like Poriferans, they have two germ layers – but these are two true tissues (endoderm/gastroderm and the ectoderm). • They also have a mesoglea – which is a jelly-like layer between the germ layers. • They have a nerve net and coordinated muscular movement– which means they can respond to their environment. • They also have specialized digestive tissues. Examples • This phylum is exclusively aquatic. • Most of the 10,000 species are marine. • Class Hydrozoa – Hydra (Freshwater) - Portuguese Man Of War • Class Scyphozoa - Jellyfish • Class Anthozoa – Sea Anemones - Coral Man ‘o’ war Feeding • They have special stinging cells called Cnidocytes. • These contain coiled toxic harpoon-containing capsules called Nematocysts. • There are thousands of these in one tentacle of an anemone, for example. • Once the food is debilitated it can be brought to the twoway mouth/anus. http://www.youtube.com/watch?v= Kukv0AtIVdU&feature=related Alternation of Body Forms - Morphology • Two main body plans are seen • Polyp and Medusa • The polyp has a sessile, tube shaped form, with the mouth located at the top. • The medusa is bell shaped and has a mouth on the underside – jellyfish. • Not all species of cnidarians have both forms. The Hydra was a many-headed monster slain by Heracles (AKA Hercules). As one of his Labors, Heracles sought the Hydra's lair in the swamps of Lerna and forced it out into the open with flaming arrows. Wading bravely into the fray, he began to hack at the monster with his sword. But every time he cut off one head, two grew in its place. Eventually, Heracles called on his charioteer to bring a torch to cauterize the Hydra's severed neck each time a head was lopped. This prevented new heads from sprouting. And when the final head was chopped off and buried beneath a rock, the monster died. Medusa was a terrible monster who had laid waste to the country. She was once a beautiful maiden whose hair was her chief glory, but as she dared to vie in beauty with Athena, the goddess deprived her of her charms and changed her beautiful ringlets into hissing serpents. She became a cruel monster of so frightening an aspect that no living thing could behold her without being turned into stone. All around the cavern where she dwelt might be seen the stony figures of men and animals which had chanced to catch a glimpse of her and had been petrified with the sight Alternation of Body Forms • The medusa reproduce sexually. • They have interesting semi-permanent gonads – cells migrate to one spot to produce gametes. • Upon fertilization the zygote grows into a ciliated larva – planula. • The planula attaches to the bottom and begins the polyp stage. • See page 567 Alternation of Generations • Hydra do not have a medusoid stage. • The reproduce asexually by branching or budding. • Some hydra can reproduce sexually because they are hermaphrodites. http://www.youtube.co m/watch?v=489CSop00 sY&feature=related Colonies of Cnidarians • Corals especially, can form very complex colonies. • The diversity of corals in the ocean is amazing, and each different type is made up of thousands of tiny individual animals. • Obelia, is a colonial Hydrozoan. Ecological Role • Symbiosis – anemones and clown fish • Important food source for sea turtles, fish etc. • Examples: • Box jellyfish – up to 3 m tentacles, kill a grown man. • Man’o’war