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1 General Characteristics • They are radially symmetrical; oral end terminates in a mouth surrounded by tentacles. • They have 2 tissue layers • Outer layer of cells - the epidermis • Inner gastrodermis, which lines the gut cavity or gastrovascular cavity (gastrodermis secretes digestive juices into the gastrovascular cavity) 2 General Characteristics In between these tissue layers is a noncellular jelly-like material called mesoglea Manubrium – tubelike structure that hangs from the medusa’s oral surface (mouth is found at the end). 3 Body Structure Hydrostatic skeleton – water or body fluids confined in a cavity of the body and against which contractile elements of the body wall act. Statocyst – a small sac surrounding a calcium carbonate concretion. “Stone” moves in response to the pull of gravity, initiating nerve impulses. Oral disc – area surrounding mouth. Contains openings which permit water to circulate. Acontia – “Threads” found at the end of mesenterial filaments which contain cnidocytes. Subdue live prey in the gastrovascular cavity. 4 Cnidarian Body Plan Polyp Form Medusa Form • Tubular body, with the mouth directed Bell-shaped or umbrella upward. • Around the mouth are a whorl of feeding tentacles. • Only have a small amount of mesoglea • Sessile shaped body, with the mouth is directed downward • Small tentacles, directed downward. • Possess a large amount of mesoglea • Motile, move by weak contractions of body 5 Cnidarian Body Plan 6 Tentacles Prey capture is enhanced by use of specialized stinging cells called cnidocytes located in the outer epidermis. Each cnidocyte has a modified cilium - cnidocil, and is armed with a stinging structure called a nematocyst. The undischarged nematocyst is composed of a long coiled thread 7 Tentacles When triggered to release, either by touch or chemosensation, the nematocyst is released from the cnidocyte and the coiled thread is everted Some nematocysts function to entangle the prey; others harpoon prey and inject a paralyzing toxin 8 Cnidarian Life Cycle Stage 1 – Adult Medusa Stage 2 – Planula larva Stage 3 - Polyp9 Reproduction • One of the most amazing adaptations is the ability of some • • • • cnidarians to regenerate lost parts or even a complete body Asexual reproduction is common with new individuals being produced by budding Sea anenomes engage in a form of asexual reproduction called pedal laceration Cnidariand are dioecious Fertilization is external, with the zygote becoming a elongated, ciliated, radially symmetrical larva - planula larva 10 Nutrition Cnidarians are carnivores with hydras and corals consuming plankton and some of the sea anenomes consuming small fishes They use they tentacles to capture prey and direct it toward the mouth so that it can be digested in the gastrovascular cavity via secretions from gland cells (extracellular digestion); some food is phagocytized by special cells and digestion occurs intracellularly The gastrovascular cavity exists as 1 opening for food intake and the elimination of waste There is no system of internal transport, gas exchange or excretion; all these processes take place via diffusion 11 Movement The cnidarian body is capable of some kind of coordinated movement Both the epidermis and the gastrodermis possess nerve cells arranged in a loose network - nerve net (plexus), which innervate primitively developed muscle fibers that extend from the epidermal and gastrodermal cells Stimulus in one part will spread across the whole body via the network 12 Response to Stimuli No cephalization or nervous system Nerve net around mouth Sensory Cells Chemoreceptors (chemicals) Thigmoreceptors (touch) Photoreceptors (light) Ocelli (eyespots) Statocysts (balance) 13 Class Hydrozoa Includes the solitary freshwater hydra; most are colonial and marine Typical life cycle includes both asexual polyps and sexual medusa stages; however, freshwater hydras and some marine hydroids do not have a medusa stage 14 Solitary Hydras Freshwater hydras are found in ponds and streams occurring on the underside of vegetation Most possess a pedal disc for movement, mouth, hypostome surrounded by 6-10 tenetacles Mouth opens to the gastrovascular cavity The life cycle is simple: eggs and sperm are shed into the water and form fertilized eggs; planula is by passed with eggs hatching into young hydras Asexual reproduction via budding 15 Colonial Hydrozoans • Possess a skeleton of chiton that is secreted by the epidermis • All polyps in the colony are usually interconnected • Two different kinds of individuals that comprise the colony: feeding polyps or gastrozooids (C) and reproductive polyps or gonozooids (B) 16 Other Hydrozoans Portuguese man-of-war: Single gas-filled float with tentacles Tentacles house the polyps and modified medusae of the colony http://www.youtube.com/ watch?v=xTgLTbXJrfM 17 Class Scyphozoa True Jellyfish The medusae are large and contain massive amounts of mesoglea The differ from the hydrozoan medusa in that the lack a velum Possess four gastric pouches lined with nematocysts; these are connected with the mouth an the gastrovascular system http://www.youtube.com/watc h?v=u0I-3wkH37w 18 Class Anthozoa: Corals • Exclusively marine; there is no medusa stage • At one or both ends of the mouth is a ciliated groove called the siphonoglyph; generates a water current and brings food to the gastrovascular cavity • Possess a well developed pharynx • The gastrovascular cavity is large and petitioned by septa or mesenteries; increase surface area for digestion or support • Edges of the septa usually have threadlike acontia threads, equipped with nematocysts and gland cells 19 Coral Video http://www.youtube.com/watch?v=-O-m_XIrA1k 20 Class Anthozoa 21 Class Anthozoa • Solitary anthozoans include sea anemones • Most anthozoans are colonial (e.g. corals) and secrete external skeletons composed of calcium carbonate. • Corals obtain much of their energy from microscopic photosynthetic green algae (zooxanthellae) or dinoflagellates that live symbiotically inside the cells of the coral 22 Phylim Ctenaphora Contains Comb Jellies Characteristics of Ctenophora: Biradial symmetry Monoecious Gastrovascular cavity Rows of ciliated bands, called “comb rows” No nematocysts; Adhesive structures known as colloblasts http://www.youtube.com/watch?v=39qe0E ajbIA 23 Comb Jellies 24