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I. Platyhelminthes • • Flatworms - Dorsoventrally flattened Simplest bilaterally symmetrical organisms • • First organs and organ systems** Central nervous system • • Simple “brain” coordinates muscle movements Incomplete digestive system • • • Mouth but no anus Similar to Cnidaria and Ctenophora Mesoderm** • • Cell layer between endoderm and ectoderm Gives rise to muscles, reproductive system I. Platyhelminthes A. Turbellaria • • • Mostly free-living carnivorous species Most commonly seen (Why?) Some live as commensal animals inside other invertebrates (oysters, crabs, etc.) I. Platyhelminthes B. Trematoda (Flukes) • Most species (6000) • • Complex life cycles • • • C. Parasitic – Feed on tissues, blood, gut contents Adults live in vertebrate host (fish, whale, bird) Larvae may inhabit invertebrates (intermediate hosts) Vertebrate eats intermediate host (clam, snail, etc.) Cestoda (Tapeworms) • • Parasitic Live in vertebrate intestines • • • Head attaches to intestine wall with suckers or hooks Gutless – absorb nutrients through body wall May reach 50 feet!! (sperm whales) II. Nemertea • • Ribbon worms Cosmopolitan • • Complete digestive tract** • • • Mouth and anus Circulatory system** Use long, fleshy proboscis to capture prey • • • Most common in shallow, temperate regions Feed on crustaceans and worms May be cryptic or conspicuous Can reach 30 m in length!! III. Lophophorates • Three phyla – all animals possess lophophore • • Ciliated hollow tentacles arranged in a horseshoe Suspension feeders • Bilateral symmetry, coelom (body cavity), U-shaped gut Ectoprocta – Bryozoans A. • Colonies consist of interconnected individual zooids • • Encrusting and lacy forms (CaCO3 tests) Retractable lophophore Phoronida – Phoronids, Horseshoe Worms B. • • Worm-shaped Agglutinated sediment tubes attached to hard substrate in shallow water Brachiopoda – Lamp Shells C. • • • Abundant in fossil record Superficially resemble clams, but shells are dorsal-ventral, not left-right as in mollusks Many attached to substrate with pedicle (short stalk) Bryozoans III. Lophophorates • Three phyla – all animals possess lophophore • • Ciliated hollow tentacles arranged in a horseshoe Suspension feeders • Bilateral symmetry, coelom (body cavity), U-shaped gut Ectoprocta – Bryozoans A. • Colonies consist of interconnected individual zooids • • Encrusting and lacy forms (CaCO3 tests) Retractable lophophore Phoronida – Phoronids, Horseshoe Worms B. • • Worm-shaped Agglutinated sediment tubes attached to hard substrate in shallow water Brachiopoda – Lamp Shells C. • • • Abundant in fossil record Superficially resemble clams, but shells are dorsal-ventral, not left-right as in mollusks Many attached to substrate with pedicle (short stalk) III. Lophophorates • Three phyla – all animals possess lophophore • • Ciliated hollow tentacles arranged in a horseshoe Suspension feeders • Bilateral symmetry, coelom (body cavity), U-shaped gut Ectoprocta – Bryozoans A. • Colonies consist of interconnected individual zooids • • Encrusting and lacy forms (CaCO3 tests) Retractable lophophore Phoronida – Phoronids, Horseshoe Worms B. • • Worm-shaped Agglutinated sediment tubes attached to hard substrate in shallow water Brachiopoda – Lamp Shells C. • • • Abundant in fossil record Superficially resemble clams, but shells are dorsal-ventral, not left-right as in mollusks Many attached to substrate with pedicle (short stalk) IV. Corals A. Biology • • • Phylum Cnidaria, Class Anthozoa (most) Lack medusa stage Hermatypic (reef building) corals produce skeletons made of calcium carbonate • Most contain mutualistic zooxanthellae (provide nutrition; enhance calcium carbonate deposition) • Many growth forms IV. Corals A. Biology • Reproduction • • • Sexual: Most broadcast spawners; some brooders • Polyp Planula (planktonic larva) Polyp Asexual: Fragmentation (can be *very* important) All polyps in a colony genetically identical Fig. 15-4 V. Coral Reefs A. Distribution • Living reefs ~ 600,000 km2 (0.17% of sea floor) Great Barrier Reef = Largest • • • • >2000 km long Up to 145 km wide Visible from space V. Coral Reefs B. Conditions 1. Substrate • 2. Primarily in areas with hard substrate Light - Bright (Why?) • • 3. Rarely develop in water > 50 m deep Continental shelves, around islands, tops of seamounts Temperature • • Mean annual water temperature > 20 oC Best development at 23-25 oC Distribution of Coral Reefs Fig. 15-14 V. Coral Reefs B. Conditions 3. Temperature • • • Fig. 15-33 Too warm also problematic • Can cause bleaching and eventually death Bleaching events often occur during periods of unusually warm water • Extreme low tide • El Niño event Corals live near their upper thermal tolerance levels • Corals from warmer waters have higher thermal tolerance levels V. Coral Reefs B. Conditions 4. Salinity • 5. Reefs tend to be absent or poorly developed near mouths of rivers Sediments • • 6. Turbidity reduces light levels Sediments can smother corals Pollution • • 7. Corals sensitive to pesticides and other chemicals Fertilizers support growth of algae that smother corals • Most corals grow in areas with low nutrient levels Tidal Regime • Most corals intolerant of prolonged exposure