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http://www.palaeos.com/Invertebrates/Molluscs/Images/mollusc_classes.jpg Phylum Mollusca • Triploblastic- 1st with coelom (small cavity space around various organs- for molluscsheart and gonads) • Bilateral symmetry (3 body parts: head-foot, visceral mass, and mantle) • Visceral mass contains all organs • Mantle is outer layer and may secrete a shell • Mouth has radula structure which is a belt with rows of curved teeth on a tongue-like structure called an odontophore. http://www.science.mcmaster.ca/Biology/Harbour/SPECIES/BIVALVIA/IMAGES/ORGAN1.GIF http://upload.wikimedia.org/wikipedia/commons/thumb/4/45/Radula_diagram3.png/300px-Radula_diagram3.png Radula and odontophore Molluscan shell loss and reduction… http://images.google.com/imgres?imgurl=http://www.andrewgray.com/essays/cepshell.gif&imgrefurl=http://www.andrewgray.com /essays/molluscs.htm&usg=__pHYWvKpK6tG9esL_b7xpQx8mQ8w=&h=234&w=312&sz=7&hl=en&start=2&um=1&tbnid=S2drmzyr1D DJZM:&tbnh=88&tbnw=117&prev=/images%3Fq%3Dmollusc%2Bloss%2Bof%2Bshell%26hl%3Den%26um%3D1 Class Gastropoda • Largest class of mollusks! (Snails, slugs, and limpets) • Undergo torsion in early development (body twisting 180°) – 3 advantages to torsion: • Head enters shell first to escape predators • Anterior opening allows clean water to enter mantle • Sensory organ around head help snail to be more sensitive to stimuli *some snail go through detorsion (twist 180then untwist 90) • • • • • • • • • • Most shells asymmetrical Feed by scraping with radula Cilia in digestive tract as well as on foot Some gastropods have a siphon =rolled extension of the mantle Respiration is by gills in aquatic snails, and by gas exchange through mantle in land snails Open circulatory system with heart Hydraulic skeleton of blood Well developed sensory structures (tentacles with eyes) Most marine snails dioecious, others monoecious Eggs laid in external fertilization, deposited in moist environments http://www.marietta.edu/~biol/ biomes/images/shores/shield_li mpet_6150.jpg Limpets http://www.guardianseries.co.uk/resources/images/23155 4/?type=display http://oceanexplorer.noaa.gov/explorations/0 4fire/logs/april14/media/limpet_eggs_600.jpg Barnacles- NOT mollusks! http://www.mtkembla-e.schools.nsw.edu.au/gallery/galleryimages/barnacles.jpg http://www.restaurantwidow.com/images/20 07/04/27/barnacles.jpg Class Bivalvia • 2nd largest mollusk class (clams, oysters, mussels, scallops) • 2 halves = valves • Oldest part of shell = umbo (near anterior) • Hinges of protein to keep shell closed • Pearls made from secretions of mantle (best pearls come from pacific oysters) • Loss of head and radula • Folded Gills for resipration • Filter feed from water: cilia in bivalve moves food along food grooves toward mouth; labial palps sort food • Open circulatory system • Nervous system with ganglia (only some species have eyes- scallops) • Most dioecious, few monoecious • External fertilization- most brood young http://oregonstate.edu/dept/ncs/photos/oysters.jpg http://blueoceannotes.files.word press.com/2009/08/oysters.jpg http://www.pac.dfo-mpo.gc.ca/science/speciesespeces/shellfish-coquillages/geopath/Images/intro1.jpg http://www.barrierreefaustralia.com/IMAGEGALLERY/giant-clam.jpg http://community.adn.com/sites/community.adn.com/files/i mages/Weathervane%20scallops%20-%20ASMI.preview.jpg http://cookingoysters.co.uk/wp-admin/images/preparing%20mussels.jpg http://wednesdaychef.typepa d.com/photos/uncategorized/ mussels_1.jpg http://oceangrow.septentriones.com/data/upi mages/scallop.jpg Class Cephalopoda • Octopus, squid, cuttlefish, nautilus • Most complex of phylum mollusca • Foot modified into tentacles/arms and funnel for locomotion • Head is in line with visceral mass • Mantle has radial and circular muscles • Possess jaws and radula (octopus inject venom) • Locate prey by sight and capture with tentacles (mostly fish, crustaceans, and other invertebrates) • Nautilus is the only cephalopod with an external shell • Cuttlefish and squid have internal skeletons – Cuttlefish = Cuttlebone – Squid = Pen • The octopus has no shell • Closed circulatory system • Pigment cells (chromatophores) to change color • Complex nervous system- large brain/ganglia – Eyes similar to vertebrates • Ink glands behind anus • Dioecious • Reproduction by male placing spermatophore into female near oviduct; eggs are laid singly or in strings and usually attach to substrate. Cuttlefish http://www.oceanfutures.org/spongebob/gallery/images/gallery_pim432c3.jpg http://fundiesvatheists.lefora.com/composition/atta chment/c441ea1fa6c330e0c9bc72ff5c5e7d54/19297 0/ComCuttlefish9.jpg Squid http://images.google.com/imgres?imgurl=http://nea torama.cachefly.net/images/2008-04/giantsquid.jpg&imgrefurl=http://www.neatorama.com/20 08/04/30/colossal-squid-at-te-papa-museum-in-newzealand/&usg=__1XokKm5sRqqkWnKCQx86OvA5f8I= &h=374&w=500&sz=46&hl=en&start=31&um=1&tbn id=qXUY7pVNM1cWgM:&tbnh=97&tbnw=130&prev =/images%3Fq%3Dsquid%26ndsp%3D20%26hl%3De n%26sa%3DN%26start%3D20%26um%3D1 http://images.google.com/imgres?imgurl=http://news.nationalgeographic.com/news/2007/02/images/070222squid_big.jpg&imgrefurl=http://news.nationalgeographic.com/news/2007/02/070222-squidpictures.html&usg=__b5FBOMpbizi4ZbuOxcWpzGr9uRI=&h=454&w=461&sz=59&hl=en&start=28&um=1&tbnid=1G9OtH5 NzdUoQM:&tbnh=126&tbnw=128&prev=/images%3Fq%3Dsquid%26ndsp%3D20%26hl%3Den%26sa%3DN%26start%3D20% 26um%3D1 http://www.scuba-equipmentusa.com/marine/FEB04/images/Octopus_tetricus.jpg Octopus http://www.smh.com.au/ffximag e/2005/02/12/octopus_wideweb __430x308.jpg http://media.smithsonianmag.com/images/deep4.jpg http://www.bbc.co.uk/devon/content/images/2007/12/06/blue_ringed_octopus_465x310.jpg