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Ch 27: Molluscs and Annelids There are some things on the review sheet and test that you should add to this presentation – eg – mussels have special glue Phylum Mollusca Molluscs – Characteristics that Unify Have true coelom Are protostomes Have trochophore larvae (like annelids) Have foot, mantle, visceral mass, and possibly a shell Protostomes, Coelomates Trochophore larvae Molluscs and Annelids – a common ancestor? Internal Structures of Molluscs Foot – structure involved in feeding, usually contains the head as well Visceral mass – contains the internal organs Mantle – secretes the shell Shell - protection Molluscs – classified by the modifications of the Foot Gastropods – “stomach foot” Foot is broad and muscular found along the ventral surface used for moving, eating Examples: Snails, slugs, nudibranchs Class Gastropoda - Snails Snail Diagram Gastropods - Sea slugs/nudibranchs Cephalopod – the tentacled molluscs “Head –foot” Foot is modified into head with tentacles coming out of it Have very little shell if any Ex. Octopi, squid, nautilus, cuttlefish, Cephalopod – the tentacled molluscs Cephalopods have a ‘head’ Cephalopods include: Octopi Squid Cuttlefish Nautiluses Have internal shells Swim with jet propulsion Bivalves Have 2 shells – foot can extend out of shell Clams, mussels, oysters Class Bivalvia 2 shells Clams Oysters Mussels Typical Shell/Valve Feeding - Molluscs Many molluscs have a structure that is like a very rough tongue – has hundreds of tiny teeth for grinding through shells or scraping algae off of rocks Molluscs – Filter feeders filter feeders use their GILLS to catch the tiny food particles They also use the gills to BREATHE Filter Feeders – Environmental “Watchdogs” Filter feeders remove the pollutants from the water and concentrate them in their bodies Environmentalists can use these animals to learn information about levels of pollution in the water. Don’t eat bivalves during a RED TIDE Red Tide Clam – How it eats Incurrent siphon pulls water across gills Food particles stick to gills Coordinated cilia move food to mouth Food is stored/digested in stomach and intestines Solid wastes exit via the anus Respiration - Molluscs Aquatic molluscs use gills Land snails and slugs use a moist sac inside the mantle Sac’s lining is convoluted (to increase surface area) and lined with blood vessels Molluscs Circulatory Systems Marine snails, slugs, clams: Slow-moving molluscs have “open” circulatory systems (like some insects) Blood vessels connect to sinuses where oxygen transfer takes place Octopi, squid: Fast-moving molluscs have “closed” circulatory systems – blood is completely enclosed in blood vessels (like ours) Circulatory System Comparisons Excretion – Molluscs and Annelids Molluscs and annelids are coelomates and protostomes – that means they definitely have an anus for solid wastes! - they use NEPHRIDIA for metabolic wastes (like the nephrons in our kidneys Nervous Systems -Molluscs Slow moving molluscs have simple systems with simple sensory cells like the cnidarians and the worms – statocysts, ocelli, chemoreceptors, touch receptors, etc. Scallop Nervous Systems - Molluscs Fast-moving molluscs have very complex nervous systems Octopi: Well-developed brain (with memory) Eyes that are similar to ours – form images Reproduction - Molluscs MOST have separate sexes that employ external fertilization (fertilization by chance!) Tentacled molluscs and some snails have internal fertilization Some are hermaphrodites Molluscs - reproduction Oysters – hermaphrodites that change sexes each season – sometimes they are the males, sometimes the females. Defence Mechanisms Gastropods Slugs hide under rocks or only come out at night Some gastropods are poisonous – brightly colored to warn off predators Some nudibranchs save the nematocysts from the cnidarians they ate, and sting their predators with them Poisonous ! Cephalopods – Defence Mechanisms Octopi move rapidly backwards with their jet propulsion of water Some release foul-tasting black ink Some can change colours Squid – 8 arms + 2 long tentacles Annelids – Characteristics that unify Segmented worms Coelomates Trochophore larvae protostome Segmentation Annelids – Feeding Filter feeders – use gills to capture food particles from water Others use pharynx to either suck in food or to extend out and get it. Some annelids have jaws on their pharnyx Christmas tree worm Nereis – (clam worm) A Polychaete – has jaws for eating and fighting Respiration Earthworm cuticle Aquatic annelids use gills Terrestrial annelids exchange gases through their skin – their skin must remain moist for this purpose Cuticle on the earthworm helps prevent desiccation Feather-duster worm – gills exposed (red) - tentacles for gathering food Internal Transport Closed circulatory system Dorsal surface has blood moving toward the head, ventral surface moves blood away from the head Each segment has a ring vessel connecting the ventral with the dorsal Large ring vessels at anterior end act as a “heart” to pump the blood If no heart is present, the bodies muscular contractions move the blood around Excretion Solid wastes out the anus Metabolic (liquid) wastes are eliminated by nephridia (like the nephrons in our kidneys) 2 nephridia/segment Response Brain on dorsal surface Nerves travel from the brain, around the gut and a main nerve cord travels down the ventral surface Brain = #11 Ganglia = #12 Ventral Nerve cord = #13 Sense organs Spaghetti worm underground With tentacles exposed Best developed in free living annelids Most of these have sensory tentacles, statocysts, chemoreceptors, and “eyes” Some form images, some are light-detectors Some have sensory organs that detect vibrations (like our tympanic membranes) Self Defence Tube worms Burrow into ground or swim away Setae (oneway bristles) anchor worms in the soil Marine fan worms make tubes of CaCO3 to hide in Self-Defence Marine fireworms have poisonous bristles (burns) Carnivorous annelids have jaws to bite their predators Movement longitudinal muscles shorten the body Circular muscles contract to decrease the diameter of the worm Coordination of these sets of muscles allow the worms to wiggle, burrow and swim Reproduction Some reproduce by budding Some have separate sexes and have external fertilization Palolo worms congregate at the surface and release sperm or eggs at the same time Feast for the predators (humans included!) Earthworms are hermaphrodites and exchange sperm Sperm is stored in sacs until eggs are ready Mucus ring is secreted by the clitellum – sperm and eggs are deposited there and the ring slips off the worm, forming a cocoon for the fertilized eggs Common Characteristics in all Annelids Unidirectional Digestive System Closed Circulatory System First sign of a ‘heart-like’ structure Nephridia used for excretion Primitive Brains present Class Polychaeta – bristleworms Mainly marine worm Have padddle-like appendages with bristles Show cephalization More Polychaetes Polychaetes Class Oligochaeta Earthworms and Relatives Earthworms + Freshwater worms Aquatic – eg tubifex (red, thread-like worms sold for fish food) Oligo – means few (as in few bristles) Castings are excellent fertilizer Class Hirudinea Commonly known as the LEECHES Fresh and saltwater species (~300) Few land species Muscular proboscis is forced into skin Others slice skin with their jaws Leeches Hirudin – anticoagulating chemical is used to keep host’s blood flowing Some also use an aesthetic so the host does not know they are there Used for medicinal purposes Annelids aerate the soil by burrowing Return nutrients to the soil completing decomposition of material Many annelids are the basis of food chains Internal Anatomy Identifying practice: