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What is a scientific theory? – It makes falsifiable predictions with consistent accuracy across a broad area of scientific inquiry – It is well-supported by many independent strands of evidence, rather than a single foundation – It is consistent with pre-existing theories and other experimental results – It can be adapted and modified to account for new evidence as it is discovered, thus increasing its predictive capability over time. – It is among the most parsimonious explanations, sparing in proposed entities or explanations. Review • Macroevolution – If the changes are over a very long time and are large enough that the population is no longer able to breed with other populations of the original species, it is considered a different species. • Microevolution – changes to the varieties of individuals in a population such as a change in a species' coloring or size. • Gene Pool – all genes in a population • Mutation – – random change in structure of DNA. Every so often, a mutation is beneficial for survival. Review • Natural Selection – a) Genetic variability in a trait within population b) Trait is heritable c) Differential reproduction – must enable individuals with the trait to leave more offspring than others without the trait. Review • Sexual Dimorphism: – The males & the females of a species have a noticeable physical difference Review • Speciation – – The formation of a new species from an ancestral species • Geographic Isolation – – Species are unable to mate/blend their genes due to the location(s) of species within a habitat/globally • Reproductive Isolation – – Behavioral or physiological barriers that prevent the reproduction between species. Review • Background Extinction – – A slow rate of die off, often due to natural selection • Mass Extinction – – A quick & massive die off of an entire population or many populations • Adaptive Radiation – – after mass extinctions, numerous new species evolve to fill vacated niches. Takes 1-10 million years for adaptive radiation to rebuild biodiversity. Review • Artificial Selection – – artificially selecting superior genetic traits – Agriculture, Hatcheries, Pets • Genetic Engineering – • Gene splicing • Species creation in laboratories • Takes less time than artificial selection • Age of the Earth – – 4.57 billion years old Review • Eon Era Period Epoch • Cambrian Period – – Sudden appearance in the fossil record of many new phyla (Cambrian explosion) – Abundant marine invertebrates • Ordovician Period – – Earliest vertebrates appeared (ostracoderms) – The first plants invaded land Review • Silurian Period – – Coral reefs became abundant – First jawed fish appeared (Placoderms) • Devonian Period – – First fish evolved into tetrapods and invaded land – Cartilaginous fish (sharks) became dominant • Carboniferous Period – – The first reptiles evolved from amphibians Review • Permian Period – – Reptiles thrived and began their radiation – Ended with the largest extinction event on record – 96% of animal species disappeared • Triassic Period – – Reptiles returned to the seas – Beginning of the radiation of dinosaurs • Jurassic Period – – Warm climate and high sea levels – First small mammals appeared – First evidence of feathered birds (Archaeopteryx) Review • Cretaceous Period – – Mammals evolved into three forms (monotreme, marsupial, placental) – all dinosaurs and all ammonites, along with many other species, became extinct • Tertiary Period – – primitive apes; first elephants – First ancient hominids arose Review • Quaternary Period – – Pleistocene: modern humans evolved; ended with the Ice Age that saw the extinction of mammoths and other animals – Holocene: human civilization spread throughout the globe and humans became the dominant form of life Review • Tiktaalik roseae – important transitional fossil discovered in 2005 that connects aquatic species to terrestrial species Review • Changes in the skull of tetrapods: – Skull becomes flattened – Eyes move from the side of the head to the top and to the front – Pectoral girdle (bones around the neck) are reduced so that the head can turn – Snouts/jaws become elongated – Primary palate expands Review • Terrestrialization (33 & 34): – Bones become more dense to support weight without the buoyant force of water – Fin rays become digits – organization of myomeres (striated muscles) in fishes (locomotion) – Lungs & pump (diaphragm, buccal pump, etc.) – Thickening of dermis – Body temperature control Review • Aquatic Mammal Examples: – Cetaceans, Sirenias, Pinnipeds • Aquatic/Terrestrial Mammal Similarities: – Endotherms – Placental (belly buttons) – Mothers produce milk – Most have hair at some stage of their development Review • Aquatic/Terrestrial Mammal Differences: – Modified limbs into flippers – Some have reduced pelvises & femurs – Cetaceans have flukes – Cetaceans have nostrils located on the dorsal portion of their head (blowhole) Review • Vestigial Features: – Genetically determined structures that have lost their ancestral function – Examples: • Femur bones & reduced pelvises in dolphins & whales • Olfactory nodes found in the brains of most cetaceans • Homologous bone structure • Atavism: – Reverting (genetic error of expression) to an ancestral type – Examples: Dolphin legs, human tails, chicken teeth Review • Conditions that favor fossilization – – Having Hard parts – shells, bones, cysts – Get buried, trapped • Marine species • Marsh, flooding areas – Abundant species (with many individuals) – Long lived species (as a species) – Avoid eroding away Review • Homologous Structures – – Similarities in body structures due to common ancestry • Analogous Structures – – Structures with similar function that do NOT share common ancestry Review • Population Genetics – – The study of genetic changes in populations over time – Quantify molecular differences within and among populations • Traits exhibit variation • Gene Flow – – The loss or gain of alleles (genes) in a population due to migration of fertile individuals or gametes between populations a)Immigration b) Emmigration Review • Genetic Drift – – a change in allele frequency due to random chance – Affect small populations – Limits alleles available to subsequent generations • Founder Effect – – Genetic drift attributed to colonization by a limited number of individuals from a parent population • Bottleneck Effect – • Genetic drift resulting from the reduction of a population, typically • by natural disaster, surviving population does not represent the original • population’s gene pool Review • Directional Selection – – 1 extreme is favorable • Diversifying Selection – – 2 extremes favorable • Stabilizing Selection – – Intermediate favored Review • Pre-zygotic Barriers – – prevent successful fertilization between species • Post-zygotic Barriers – – allow fertilization but prevent successful development / reproduction of hybrid. Review • Habitat Isolation – – Two organisms that use different habitats even in the same geographic area are unlikely to encounter each other to even attempt mating. • Behavioral Isolation – – Many species use elaborate behaviors unique to that species to attract mates. • Temporal Isolation – – Two species that breed during different times of day, different seasons, or different years cannot mix gametes. Review • Mechanical Isolation – – Physical/ structural mechanism that prevents reproduction • Reduced Hybrid Viability – – Miscarriage, early death Review • Reduced Hybrid Fertility – – Sterile – Unattractive to opposite sex • Hybrid Breakdown – – 1st generation is viable & fertile – 2nd generation is unviable or infertile – Occurs in plants • Allopathic Speciation – – A mode of speciation induced when the ancestral population becomes segregated by a geographic barrier Review • Parapatric Speciation – – Takes place in large range – Variable environmental – Variable selection pressure – Non-random mating • Sympatric Speciation – – – – – Takes place within habitat of parent species Reproductive barrier forms within a subset of population Maybe genetic that immediately blocks reproduction Maybe behavioral Review Review • Phylogeny – – Technique to understand evolutionary relationships • Phylogenetic Trees – Review • Convergent Evolution – – Evolutionary pattern where two or more non related taxonomical groups independently evolved same/similar characteristics – Similar structures evolve under similar environmental conditions