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The Origin and Early Evolution of Life Chapter 18 Learning Objectives 1. 2. 3. 4. Differentiate between chemical evolution and organic evolution. Describe the conditions on the primitive earth predicted by laboratory experiments that favored the spontaneous development of organic molecules. Explain the probable sequence in the origin of the first cells, and describe how this cell interacted with its environment. Review the theories proposed to explain the origin of eukaryotic organelles. The Big Bang • 12-15 billion years ago all matter in the universe was compressed into a mass the size of our sun • Sudden instantaneous explosion of matter and energy throughout the known universe p. 288 • About 4.6 to 4.5 billion years ago • Minerals and ice orbiting the sun started clumping together • Heavy minerals moved to Earth’s interior, lighter minerals floated to surface – Produced outer crust and inner mantle • Chemical evolution produced the primordial sea and the early atmosphere that would support the biosphere of the future. Earth Forms Earth Is “Just Right” for Life The Planet: Early Atmosphere: • If smaller in diameter, gravity would not be great enough to hold onto atmosphere • Hydrogen gas • If closer to sun, water would have evaporated • If farther from sun, water (the solvent of life) would have been locked up as ice • Nitrogen • Carbon monoxide • Carbon dioxide • No gaseous oxygen p. 288 Stanley Miller’s Experiment • Mixed methane, hydrogen, ammonia, and water • Simulated lightning • Amino acids and other small molecules formed electrodes to vacuum pump CH4 NH3 H2 O H2 spark discharge gases water out condenser water in What, at a minimum, is needed to form a cell-like structure from these molecules? p. 289 water droplets boiling water water containing organic compounds liquid water in trap Possible Sequence p. 291 Why is RNA believed to be the first genetic information? Why is DNA a better molecule for heredity? START: membrane-bound proto-cells living cells END “Organic Evolution” self-replicating system enclosed in a selectively permeable, protective lipid sphere DNA RNA formation of protein-RNA systems, evolution of DNA enzymes and other proteins formation of lipid spheres spontaneous formation of lipids, carbohydrates, amino acids, proteins, nucleotides under abiotic conditions informational molecules separation of environments “Chemical Evolution” The First Cells • Prokaryotic heterotrophs Describe what this means. • Secured energy through anaerobic metabolism …Describe. – No oxygen present …Why not? Is there fossil evidence for these first cells? p. 291 Fossil Evidence • photosynthetic eubacteria • Oxygen producing • permits aerobic respiration 3.5 bya p. 292 2.1 bya 1.2 bya Possible Origin of ER & Nuclear Envelope DNA infolding of plasma membrane aka: Autogenous Theory p. 294 Endosymbiotic Theory p. 295 Cyanophora paradoxa