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Unit: Plant Nutrition Nutritional Needs of Plants Plants and mineral nutrients: Plants require certain elements in order to grow and stay healthy. 16 Elements essential to plant growth: Macronutrients: • Carbon (C) • Hydrogen (H) • Oxygen (O) • Nitrogen (N) • Phosphorous (P) • Potassium (K) • Sulfur (S) • Calcium (Ca) • Magnesium (Mg) • • • • • • Micronutrients: Chlorine (Cl) Iron (Fe) Manganese (Mn) Boron(B) Zinc (Z) Copper (Cu) • Molybdenum (Mo) What makes an element essential? There are 3 established criteria for an “essential element”: 1. The plant cannot complete it’s life cycle without the element. 2. Action of the element must be specific- no other element can substitute for it. 3. The element must be directly involved in the nutrition of the plant. Macro vs. Micro • Macronutrients: nutrients required in large amounts and absorbed from the soil or hydroponic solution. • Micronutrients: nutrients required in small amounts and absorbed from a soil or a hydroponic solution. • Does NOT imply importance. All of the essential nutrients are required for growth and function. Mobile vs. Immobile These nutrients are further divided into the mobile and immobile nutrients. Mobile: A plant will always supply more nutrients to its younger leaves than its older ones, so when nutrients are mobile, the lack of nutrients is first visible on older leaves. Immobile: When a nutrient is less mobile, the younger leaves suffer because the nutrient does not move up to them but stays lower in the older leaves. Macronutrients • Carbon (atmosphere) • Hydrogen (Water) • Oxygen (atmosphere) These 3 nutrients make up 95% of a plants fresh tissue. Typically not limiting factors except for: Drought Disease Extreme cold Poor drainage N - Nitrogen • Function: Nitrogen stimulates growth, increase of fruit production and gives plants a healthy green color. • Toxicity: dark green plant with restricted root system and fruit set. • Deficiency: Mobile. Growth restricted. Leaves become light green and yellow then die. K - Potassium: •Function: Catalyst/activator for enzymes Encourages vigor/health •Deficiency: Highly mobile – older growth Poor growth, chlorosis, necrosis; reduced gas exchange •Toxicity: Hi K may cause Ca, Mg, Mn, Zn, Fe deficiencies P - Phosphorus: •Function: for the energy molecules: ATP, ADP, AMP •Deficiency: Very mobile (old growth) Lower leaf surfaces turn purple Leaves curl downward Poor leaf, root, flower development •Toxicity: Not usually a problem Ca -Calcium: •Function: As an activator of enzymes Plant strength & vigor As a “neutralizing” agent •Deficiency: Immobile (new growth) Misshapen, poor or no growth Blossom end rot of fruit •Toxicity: Not usually a problem (Mg) Magnesium: Function: “heart” of the chlorophyll molecule Enzyme activator (ATP, DNA, RNA) •Deficiency: Highly mobile Older leaf interveinal chlorosis Upturned leaves •Toxicity: Not known Sulfur: •Function: Form “disulfide bonds” that aide in the folding of proteins and effect their function •Deficiency: Moderately mobile Less chlorophyll chlorosis and purplish veins in mid/young leaves Thin, brittle stems •Toxicity: Red. Growth, interveinal chlorosis, leaf burn Let’s review the macros… (all result in poor growth) Mobile Functions Def. Symptoms N Y protein, RNA, DNA, Chl chlorosis, necrosis P Y ATP, etc. purple underleaf K Y Catalyst/activator for enzyme red. Transpiration Ca N cell walls, enzyme act. poor “new” growth blossom end rot in fruit Mg Y Chlorophyll, enzyme act. Interveinal chl. brittle upturn leaf S Mod 2 amino acids “mid/new” growth: chlorosis, purple veins, thin brittle stems Micronutrients: Iron: (Fe) •Function: Involved in chlorophyll & protein synthesis and respiration •Deficiency: Immobile New growth interveinal chlorosis •Toxicity: If over-apply foliar necrotic spots Manganese: (Mn) • Function: Enzyme activator (chl, RNA, DNA) carbohydrate metab., O2 prod. • Deficiency: Relatively immobile Newer growth interveinal chlorosis Necrotic spots/leaf drop Upward leaf curl • Toxicity: Reduced growth Hi Mn Low Fe and interveinal chlorosis Boron: (B) • Function: Related to metabolism of Ca, K Regulates carbohydrate Involved in RNA synthesis •Deficiency: Mobile in leaf; not in plant Stems: abnormal/slow growth, brittle Shoot/root die back Young leaves: thick, curled Low flowering, fruiting; fruit rots •Toxicity: Def. sym. + leaf tip chlorosis/necrosis Zinc: (Zn) • Function: Enzyme activator Synthesis of proteins, hormones, RNA, DNA •Deficiency: Not very mobile Abnormal and stunted growth Leaves: interveinal chlorosis, necrosis • Toxicity: Similar to def. symp. Copper: (Cu) • Function: (70% of Cu in leaf in chloroplasts) Chlorophyll synthesis •Deficiency: Immobile Leaf stunting, twisting; dark green Reduced turgor (firmness) Reduced flowering/fruiting • Toxicity: Do not use Cu tube/fixtures Overall stunting; thick, dark roots Molybdenum: (Mo) •Function Involved in carbohydrate metabolism. •Deficiency: Mobile Leaf interveinal chlorosis, mottling, scorching, inward cupping Severe: puffy leaf areas & stunting • Toxicity: Not usually seen… Leaves can turn yellow Chlorine: (Cl) • Function: Enzyme activator Promotes healthy growth of plants Hi NaCl stress flavor, nutrition • Deficiency: Mobile Leaf chlorosis then necrosis Low transpiration, wilting Plant stunting, die back •Toxicity: Leaf tip burn. Let’s review the micros: ** Some are mobile, some are not ** ALL are involved in plant metabolism Enzyme activators Parts of enzymes In electron transfer As oxygen carriers Invovled in synthesis/metabolism of proteins, hormones, RNA/DNA, carbohydrates, chlorophyll, ion balance, etc. ** KEY: mainly move around assisting in reactions, etc. Therefore, not needed in large amounts for “structure”. Other nutrients: Found in some but not all plants; possibly required by those plants Sodium (Na) C4 plants Silicon (Si) C4 & CAM Cobolt (Co) N-fixing bacteria Vanadium (V) Essential in green algae; toxic in hp in hi conc Iodine (I) Stimulates growth in low conc; toxic in hi conc Bromine (Br) Can substitute for Cl Fluorine (F) Toxic; some plants accumulate; hi in teas! Aluminum (Al) Req’d for tea bush; toxic for most plants Nickel (Ni) Maybe req’d for N-fixers; toxic for other plants Selinium (Se) In milk vetches; toxic to most plants Pictures from this presentation can be found at: www.plantphys.net/_article.php?ch=t&id=289 Plant Physiology online: Chapter 5.1 Visual symptoms of nutrient deficiencies in plants Dr. Wade Berry, UCLA Questions?