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CLUSIACEAE The pattern of pollen types in angiosperms Furness & Rudall, 2004 Successive sporogenesis Simultaneous sporogenesis Furness & Rudall, 2004 Furness & Rudall, 2004 eurosids I rosids magnoliids monocots core eudicots eudicots CLUSIACEAE (Clusioideae: MALPIGHIALES, EUROSID I) NUMBERS: 50 Genera, 1200 species GEOGRAPHY: pantropical ECOLOGY: small trees and stranglers of wet to rain forests mostly at middle to upper altitudes ROSID CHARACTERS: petals separate, androecium diplostemonous and/or not fused to petals MALPIGHIALES: APG SAYS: leaf margin toothed [teeth with a single vein running into a congested ± deciduous apex]. SEEMS WORTHLESS. The butterfly Cymothoë has hosts widely scattered in this order alone (Ackery 1988). CLUSIACEAE, Malpighiales Cymothoë likes Malpighiales, its allies are all Rosid specialists and mostly eurosids I specialists. CLUSIACEAE - treelets and stranglers Clusiaceae in a Golfito profile: 2, 5, 20, 24 CHARACTERS DIAGNOSTIC OF FAMILY: Habit woody Latex present - white or yellow - borne in latificers between veins of leaves Leaves opposite, secondary veins often parallel and close set Flowers imperfect (plants often dioecious, large and showy) Androecium of many showy stamens Gynoecium - styles variously connate Fruit dehiscent or not, with seeds arillate Clusia as a treelet (in montane rain forest near treeline dwarf or elfin forest Clusia as a strangler Clusia - dioecious, offers resin rewards for stingless bees. Trigona - a meliponine bee Clusia, bird-dispersed with arils as attractants. Garcinia - largely Old World, including the mangosteen Symphonia - the peppermint candy flower blue honeycreeper black-faced dacnis