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Path Of Food Through The Animal Body Chapter 24 Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Outline • • • • • Food Energy Digestive Systems Types Organs - Accessory Homeostasis Osmoregulation Vertebrate Kidney Nephrons Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Calories For Energy • Food provides animals both a source of energy and a supply of raw materials. Fats have more energy-rich carbonhydrogen bonds and thus a higher energy content than carbohydrates or proteins. Carbohydrates are obtained primarily from cereals, grains, fruits and vegetables, and contain on average 4.1 calories per gram. - Used for energy. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Calories For Energy Dietary Fats are obtained from oil margarine, and butter, and contain 9.3 calories per gram. - Used to construct cell membrane and other cellular structures. Protein can be obtained from dairy products, poultry, fish, and meats and have 4.1 calories per gram. - Used as building materials for cell structures, enzymes, hemoglobin. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Nutrition Pyramid Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Essential Substances for Growth • Many vertebrates are unable to manufacture one or more of 20 essential amino acids (necessary for metabolism), and thus must obtain them from food. Vitamins are organic substances used in trace amounts. Trace Elements are minerals required in very small amounts. - Iodine, cobalt, zinc, molybdenum, manganese, and selenium. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Types of Digestive Systems • • • Heterotrophs are divided into three main groups: Herbivores - eat exclusively plants. Carnivores - eat exclusively meat. Omnivores - eat both plants and meat. Single-celled organisms digest food intracellularly. Other animals digest food extracellularly within digestive cavity. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Types of Digestive Systems • Specialization occurs when the digestive tract (alimentary canal) has a separate mouth and anus, so that food transport is one way. Ingested food may be stored in specialized region of tract, or may first be subjected to physical fragmentation, followed by chemical digestion. Products then pass through epithelial lining of the gut into blood (absorption). Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies One-Way Digestive Tracts Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Vertebrate Digestive Systems • Vertebrate system consists of tubular gastrointestinal tract and accessory digestive organs. Mouth and Pharynx Esophagus - Delivers food to stomach. Stomach - Preliminary digestion. Small Intestine - Digestion and absorption. Large Intestine - Water/ mineral absorption. Cloaca / Rectum - Waste expulsion. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Human Digestive System Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Vertebrate Digestive Systems • In general, carnivores have relatively shorter intestines than herbivores. Plant cellulose resists digestion. - Ruminants (cows) contain fourchambered stomachs. - Other herbivores (rabbits - horses) posses cecum at beginning of large intestine. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Vertebrate Digestive Systems • Layered structure of gastrointestinal tract. Mucosa Submucosa Muscularis Serosa Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mouth and Teeth • • • Carnivorous mammals have pointed teeth, lacking flat grinding surfaces, adapted to cutting and shearing. Herbivorous mammals have large and flat teeth adapted to grinding cellulose. Human Mouths are basically carnivorous in the front, and herbivorous in the back. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mouth and Teeth • Tongue mixes food with saliva to moisten and lubricate food. Contains amylase which initiates breakdown of starch into maltose. - Presence of food in the mouth triggers increased rate of secretion, as tastesensitive neurons send impulses to the brain which responds by stimulating salivary glands. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mouth and Teeth • • Tongue moves food to the back of the mouth prior to swallowing . Soft palate elevates and pushes against back wall of pharynx. Stimulates neurons and sends impulses to swallowing center in the brain. Muscles contract and raise the larynx. - Pushes glottis against epiglottis to keep food out of respiratory tract. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Human Pharynx, Palate, and Larynx Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Esophagus and Stomach • • Swallowing center stimulates successive waves of contraction (peristalsis) that moves food along esophagus to stomach. Movement of food from esophagus into stomach controlled by either smooth muscle (sphincter) or true sphincter. Stomach is a saclike portion of digestive tract with convoluted surface enabling expansion. Allows for sporadic gorging. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Esophagus and Stomach • • Stomach contains extra layer of smooth muscle for mixing food with gastric juices. Parietal cells secrete hydrochloric acid. Chief cells secrete pepsinogen. Human stomach produces about 2 liters of HCl and other gastric secretions daily. Acidic solution kills most bacteria. - Chyme - mixture of food and gastric juices. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Esophagus and Stomach • • Hormone gastrin regulates HCl synthesis. Overproduction of gastric acid can potentially eat through stomach wall. - Gastric ulcers Susceptibility is increased when mucosal barriers are weakened by Heliobacter pylori infection. Chyme leaves stomach through pyloric sphincter and enters small intestine. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Stomach and Gastric Glands Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Small Intestine • • Small intestine breaks down large molecules into small molecules that absorb into bloodstream. Pancreas secretes enzymes into duodenum. Liver secretes bile salts into duodenum to make fats water soluble. Rest of small intestine (ilium) is devoted to absorption, and is covered with villi to increase absorptive surface. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Small Intestine Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Large Intestine • Large Intestine (colon) finishes absorption. Primary function is refuse dump. - Feces (Undigested food and bacteria) Rectum Anus Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Variations In Vertebrate Digestive Systems • Most animals lack enzymes necessary to digest cellulose. Four-chambered stomachs - Rumen - Reticulum Omasum Abomasum Coprophagy - Eating feces to absorb nutrients. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Ruminant Stomach Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Accessory Digestive Organs • Pancreas Contributes secretions to digest protein. Produces hormones in islets of Langerhans. - Controls glucose Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Accessory Digestive Organs • • Liver Largest organ in body. Delivers bile to duodenum. Modifies substances absorbed in gastrointestinal tract. - Removes toxins. Produces proteins found in blood plasma. Gallbladder Stores bile Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Homeostasis • • Dynamic constancy of internal environment. Fluctuate continuously within narrow limits. Hypothalmus coordinates body temperature regulation. Sweating, Blood Vessel Dissipation Shivering, Blood Vessel Constriction Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Homeostasis • • Islets of Langerhans secrete insulin when glucose levels rise. Liver secretes glucose when blood glucose levels fall. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Homeostasis • Catabolization produces nitrogenous wastes that must be eliminated. Ammonia Urea Uric Acid - Uricase Allantonin Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Osmoregulatory Organs • Osmoregulation is process of regulating body’s osmotic composition. Filter fluid into tubules and reabsorb ions and water. - Flatworms - Protonephridia - Earthworms - Nephridia - Insects - Malpighian Tubules - Vertebrates - Kidneys Selective absorption Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Evolution of Vertebrate Kidney • Kidney is complex organ made up of repeating disposal units (nephrons). Blood pressure forces fluid in blood past filter (glomerulus). - Retains large molecules but allows water and small molecules to pass through. Urine Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Evolution of Vertebrate Kidney • Freshwater Fish Faced two problems: - Water tends to enter body from environment. Do not drink water and excrete large volume of dilute urine. - Solutes tend to leave body and enter environment. Reabsorb ions across nephron tubules. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Vertebrate Nephron Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Evolution of Vertebrate Kidney • Marine Bony Fish Body fluids are hypotonic to seawater. - Water leaves by osmosis across gills. Drink large amounts of seawater. - Most monovalent ions actively transported out of blood across gills. - Divalent ions entering blood are secreted into nephron tubules and excreted in the urine. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Evolution of Vertebrate Kidney • • Cartilaginous Fish Reabsorb urea from nephron tubules and maintain blood urea concentration 100 times higher than that of mammals. - No net movement of water. Amphibians Produce very dilute urine and compensate for loss of sodium by actively transporting sodium across skin from surrounding water. Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Evolution of Vertebrate Kidney • • Reptiles Reabsorb salt and water in nephron tubules, and additional water in cloaca. Mammals and Birds Only vertebrates able to produce urine with higher osmotic concentration than body fluids. - Excrete wastes in small volume of water. Loop of Henle portion of nephron Cloaca in birds Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mammalian Kidney • Each kidney receives blood from renal artery and produces urine. Urine drains from each kidney through ureter which carries urine to urinary bladder. - Renal pelvis divided into: Renal cortex Renal medulla Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mammalian Urinary System Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mammalian Kidney • Nephron Regions Filter - Bowman’s Capsule Glomerulus Tube - Loop of Henle Duct - Collecting Duct Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Mammalian Urinary System Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Kidney At Work • Five Steps of Urine Formation Pressure Filtration Reabsorption of Water Selective Reabsorption Tubular Excretion Further Reabsorption of Water Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Review • • • • • Food Energy Digestive Systems Types Organs - Accessory Homeostasis Osmoregulation Vertebrate Kidney Nephrons Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies Copyright © McGraw-Hill Companies Permission required for reproduction or display Johnson - The Living World: 3rd Ed. - All Rights Reserved - McGraw Hill Companies