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Jumping, flying and swimming Movement in “fluids” Aim  jumping  gliding  powered flight  insects  birds  drag and thrust in swimming References  Schmidt - Nielsen K (1997) Animal physiology  McNeill Alexander R (1995) CD Rom How Animals move  Journals & Web links: see: http://biolpc22.york.ac.uk/404/ First: What limits jumping ? Jumping  What limits how far we can jump?  At take off have all energy stored as KE  conversion of kinetic energy to potential (gravitational) energy  KE = ½ m v2  PE = mgh How high  depends on KE at take off  PE = KE therefore mgh = ½ mv² gh = ½ v² therefore h = ½ v2/g  no effect of mass on how high you jump  neglects air resistance How far do we go?  constant acceleration due to constant gravity affected by mass  jumping in a parabola  depends on take off angle  d = (v² sin 2a) /g   jumpingangle.xls maximum at 45o  Sin 90 = 1  d = v2/g Jumping 0.12 0.1 height (m)  not 0.08 0.06 0.04 0.02 0 0  twice as far as the max height 0.05 0.1 0.15 distance (m) 0.2 0.25 0.3 How far  as before distance not affected by body mass Alice Daddy age 8 ?? mass 35kg 87kg distance 1.16m ?? Great locust jumping test  http://biolpc22.york.ac.uk/404/practicals/ locust_jump.xls Jumping in locusts  If we could jump as well, we could go over the Empire state building  max up is ½ horizontal distance  elastic energy storage  co-contraction How long to take off?  depends on leg length  time to generate force is 2s/v  for long jump, time = 2s/(g*d) s is leg length, d is distance jumped  bushbaby 0.05 to 0.1s  frog 0.06s  flea 1 ms  locust ?? Running jump  much higher/further  KE can be stored in tendons and returned during leap Summary so far  Jumping is energetically demanding  muscle mass : body mass is most important  store energy in tendons if possible Now onto: how do we fly? Flying  gliding  power flight  hovering  How stay up?  Can nature do better than mankind? Who flies?  birds  insects  bats  pterosaurs Lift  why don’t birds fall due to gravity?  where does lift come from?  speed up air  Bernoulli’s Principle  Total energy = pressure potential energy + gravitational potential energy + kinetic energy of fluid How does air speed up?  air slows down underneath because wing is an obstacle  air speeds up above wing  fixed amount of energy Lift and vortices  faster /slower airflow  =circulation  extends above / below for length of wing  creates wake Circulation  circulation vortex shed at wingtips So to fly…  we need to move through the air  use PE to glide down  as go down, PE changed to KE  use wings to force a forwards movement Can nature beat man? Gliding  soaring in thermals  Africa: thermals rise at 2-5m/s  soaring at sea/by cliffs Summary so far  Jumping is energetically demanding  muscle mass : body mass is most important  store energy in tendons if possible  Flying involves generating lift  gliding  use PE to get KE to get speed to get lift Flapping flight  large birds fly continuously  down stroke air driven down and back  up stroke  angle of attack altered  air driven down and forwards  continuous vortex wake Discontinuous lift small birds with rounded wings lift only on downstroke  vortex ring wake Summary  Jumping is energetically demanding  muscle mass : body mass is most important  store energy in tendons if possible  Birds heavier than air  Flying involves generating lift  gliding  use PE to get KE to get speed to get lift  flapping propels air Insect flight  flexibility of wings allows extra opportunities to generate lift  rotation of wing increases circulation Insect flight  flexibility of wings allows extra opportunities to generate lift  fast flight of bee  downstroke  upward  upstroke lift lift move wing bee Clap and fling  at top of upstroke two wings “fuse”  unconventional aerodynamics  extra circulation  extra force Wake capture  wings can interact with the last vortex in the wake to catch extra lift first beat second beat Summary so far  Jumping is energetically demanding  muscle mass : body mass is most important  store energy in tendons if possible  Flying involves generating lift  gliding  use PE to get KE to get speed to get lift  flapping propels air  insects often have unconventional aerodynamics – can beat the “laws” of physics  Next… Swimming Jet propulsion  conservation of momentum = m*v  mass of fish * velocity of fish = mass of water * velocity of water  squid   contract mantle dragonfly larvae Paddling / rowing  depends on conservation of momentum ducks  frogs swimming  beetles  Drag  friction  turbulence  Reynolds number gives an estimate of drag  Re = length * speed * density / viscosity  for air, density / viscosity = 7*104 s / m2  for water; density/ viscosity = 106 s/m2 Reynolds number  Re < 1 no wake  e.g. protozoan  Re < 106 flow is laminar  e.g. beetle  Re > 106 flow is turbulent  e.g. dolphin  Drag depends on shape  Drag reduced by up to 65% by mucus Design for minimal drag  tuna or swordfish:  highly efficient for high-speed cruising in calm water  torpedo-shaped body  narrow caudal peduncle  lunate, rigid fins Why don't all fish look like that?  The design is highly inefficient:  In naturally turbulent water (streams, tidal rips, etc.)  for acceleration from stationary  for turning  for moving slowly  & especially for lying still Ambush predators  keep head still  long body/dorsal fins  rapid start  flexible body, plenty of muscle  large tail fin  barracuda  pike Design for manoeuvrability  Small items don't move fast, but require delicate, focused movements for capture.  A short, rounded body with sculling or undulating fins.  Compressing the body laterally provides a wide surface to exert force on the water Optimal design?  Minimise drag often in biomechanics  No one optimal design  efficient energetics isn’t all  maximum speed isn’t all  use drag on oars to achieve efficient propulsion How does a fish move?  undulations from front to back How is thrust generated?  thrust = momentum / time  anguilliform How else is thrust generated?  tail movement  Carangiform  tail generates symmetric vortex street note rotation How else is thrust generated?  tail movement acts like a hydrofoil  thunniform  cetaceans  penguins Flying not swimming  tail movement acts like a hydrofoil  generates lift and drag  drag  lift acts in line of motion acts perpendicular (normal) to drag total lift drag Summary  Jumping is energetically demanding  store energy in tendons if possible  Flying involves generating lift  accelerate air to get lift  Insects are small enough to have unconventional aerodynamics  Minimisation of drag  Adaptation to environment leads to alternate solutions of best way to swim