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PHOTOPERIODIC INDUCTION INVOLVES PHYTOCHROME Phytochrome – pigment involved in a number of growth processes such as germination of some seeds. PHOTOPERIODIC INDUCTION Phytochrome – pigment involved in a number of growth processes such as germination of some seeds. Pr – absorbs red light at a wavelength of 660nm Pfr – absorbs far-red light at a wavelength of about 725nm. PHOTOPERIODIC INDUCTION Phytochrome – pigment involved in a number of growth processes such as germination of some seeds. Pr – absorbs red light at a wavelength of 660nm Pfr – absorbs far-red light at a wavelength of about 725nm. When Pr absorbs light it is converted to Pfr and vice versa. PHOTOPERIODIC INDUCTION Sunlight contains both red and far red light so both reactions would be expected to occur at the same time. Sunlight contains both red and far red light so both reactions would be expected to occur at the same time. However, there is more red light than far-red so red light is dominant. Most plant responses are in anticipation of environmental change as a result of slowing changing external factors – daylength. ADAPTIVE ADVANTAGES OF PHOTOPERIODISM Most plant responses are in anticipation of environmental change as a result of slowing changing external factors – daylength. Photoperiodism allows plants to: - Reach sexual maturity to coincide with pollinators being most active and present in greatest numbers increase chances of cross-pollination or reproduce asexually when conditions are optimal. ADAPTIVE ADVANTAGES OF PHOTOPERIODISM Photoperiodism allows plants to: - Expend energy and resources producing leaves etc when daylength and light intensities are at their maximum for growth and production of seeds. ADAPTIVE ADVANTAGES OF PHOTOPERIODISM Photoperiodism allows plants to: - Expend energy and resources producing leaves etc when daylength and light intensities are at their maximum for growth and production of seeds. - Drop leaves to reduce transpiration when soil water may be frozen, and develop buds that resist damage by frost/snow. ADAPTIVE ADVANTAGES OF PHOTOPERIODISM Trees in temperate climates (oak and ash) short days trigger off winter bud formation. BUD DORMANCY Trees in temperate climates (oak and ash) short days trigger off winter bud formation. These remain tiny and protected until spring – this period of dormancy can only be broken by chilling temps. BUD DORMANCY Trees in temperate climates (oak and ash) short days trigger off winter bud formation. These remain tiny and protected until spring – this period of dormancy can only be broken by chilling temps. Therefore the factor that breaks dormancy (chilling) is different from the factor that induces it (short days). BUD DORMANCY Not passive absence of growth – it is actively maintained by a growth inhibitor called abscisic acid (ABA) BUD DORMANCY Not passive absence of growth – it is actively maintained by a growth inhibitor called abscisic acid (ABA) Chilling breaks dormancy by increasing the amount of gibberellin which stimulates the growth again. BUD DORMANCY Not passive absence of growth – it is actively maintained by a growth inhibitor called abscisic acid (ABA) Chilling breaks dormancy by increasing the amount of gibberellin which stimulates the growth again. Without chilling, a plant may be late in resuming growth the next season. BUD DORMANCY Leaf fall is preceded by abscission – complex process to do with changes in the layer of cells at the base of the leaf. LEAF FALL (ABSCISSION) Leaf fall is preceded by abscission – complex process to do with changes in the layer of cells at the base of the leaf. When the leaf falls, a layer of cork seals off the leaf. LEAF FALL (ABSCISSION) Leaf fall is preceded by abscission – complex process to do with changes in the layer of cells at the base of the leaf. When the leaf falls, a layer of cork seals off the leaf. Although short days act as an external stimulus, the internal chemical signals are complex and involve interactions between several stimulus. LEAF FALL (ABSCISSION)