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"It is not the strongest or the most intelligent who will survive but those who can best manage change”- Charles Darwin. Island Geography has revealed the evolutionary process through colonization and isolation (Overview). There are fossils that date back 300 million years, providing a record of life at that time. Using these fossils is one way the theory of Island Geography is studied. Throughout this paper I will reveal the history, theories, and various concepts of Island Geography. History Biogeography dates back to the eighteenth century; starting with Georges-Louis Leclerc, Comte de Buffon. He was one of the first to question 2,000 years of the churches belief system. Unlike his colleagues, Buffon refused to accept catastrophe stories, such as the biblical flood. Sea fossils and sedimentation of rock beds indicate the continents were once submerged under water. Reemergence of them had no explanation. Buffon was an advocate of real causes and insisted these changes occurred naturally through subsidence and earthquakes. He was mostly involved with biological and zoological problems, along with key information on quadrupeds, birds, and minerals. He believed the best method was having “the complete description and exact history of each thing in particular” (ibid. p. 14B), (Buffon). “The history of one animal should be… that of the entire species of that particular animal; it ought to include their procreation, gestation period, the time of birth, number of offspring, the care given by the mother and father, their education, their instincts, their habitats, their diet, the manner in which they procure food, their habits, their wiles, their hunting methods” [ibid., 16A–B] (Buffon). Melissa Gantt Bio 111 D6 Page 1 One of the most well-known studies of Island Geography is the Galapagos Islands. From the shoreline of the equator it can take up to 600 miles to reach the cluster of islands. Because of this factor they were isolated for millions of years, creating homes for the rarest forms of plants and animals on the planet. Even now their geography is constantly changing because of the frequent volcanic activity and erosion on the Islands. It is said that the abundance of rare and unique life forms had Darwin in awe. Soon enough these incredible species created a breakthrough in his work. The way these creatures were able to flourish with such an unforgiving environment peaked curiosity; helping pave the path to Darwin’s Theory of Evolution and Natural Selection as we know it today. Many initial arrivals came to the island at its lowest. There were no mates to be greeted, no plants to be fed on, and not even soil to put down roots (Biological). Over millions of year’s natural occurrences helped populate the island, bringing birds through storms, seals and penguins through currents and carrying seeds across continents (G. islands). Melissa Gantt Bio 111 D6 Page 2 These original species could live off of what seemed to be nothing; even plants were growing out of lava. They basically survived to die; to feed the flora for the next vegetarian generation. Darwin’s interest in the island put his Theory of Natural Selection into motion. Discovery Darwin’s Theory of Natural Selection was based on what is called “Darwin’s Finches.” Originally, species were thought to be static, meaning they were thought to have no evolutionary capabilities. However, this was proven wrong by mutations in DNA. The useful genes connected to survival or breeding, help a species produce more offspring that have the same advanced gene, which will be passed down. The ones that can adapt and change will survive. Originally Darwin thought in the view of “centers of creation” he completely ignored transmutation of species. When voyaging on The Beagle they found different types of species on every island they stopped on, including 3 species of mockingbirds. Finches were also found but ignored at this point. Darwin sent his findings of birds to ornithologist expert John Gould. He discovered that the birds thought to be black birds and finches were in fact 12 species of ground finches. Every island Darwin visited had not just a variety of mockingbirds but completely different species per island. Gould found 26 species of birds, way more than Darwin anticipated, and 25 of the 26 were new and distinct species only found near the South American continent (Darwin’s). At this time the acting governor, Nicholas Lawson informed Darwin about the differences in Galapagos Tortoises and he became intrigued. Melissa Gantt Bio 111 D6 Page 3 One of the tortoises’ most unique evolutionary traits occurred because of a rice rat infestation. They ate all the food on the island and eventually tortoises evolved to be able to go without food or water for up to a year. This amazing trait indirectly led to their downfall. Buccaneers, whalers and fur sealers heard about these giant tortoises and wanted to store them on their ships for fresh meat, exploiting their adaptation (Watson). These traits made Darwin wonder about the stability of species; eventually leading to Darwin’s acceptance of transmutation of species idea. These species were similar from island to island but perfectly adapted to their environment forcing him to question the origin of the islands inhabitants (Biological). The Galapagos Islands had an enormous impact on Darwin and the world of science. This is the reason for the Charles Darwin Research Station established at Puerto Ayora, Santa Cruz Island, Galapagos in 1964 (History). The enclosure allows people to come face to face with the giant tortoises Darwin himself studied, along with an “internationally-recognized herbarium containing the largest Galapagos plant collection in the world” (History). The stations main goal is conserving life on the Islands. Unfortunately, in 2012 the last surviving Pinta Tortoise, Lonesome George, passed away. Originally they tried mating him, but nothing ever came from it. “[The Charles Darwin Foundation provides] unique scientific solutions to protect the Islands Galapagos. [Their] work continues to make the biggest impacts at local, national and international levels” (History). Melissa Gantt Bio 111 D6 Page 4 Types of Island Geography “An "island" is any area of suitable habitat surrounded by an expanse of unsuitable habitat” (Island). When mentioning islands most people think of land surrounded by water. However, there are many types of untraditional islands. Some organisms are able to live on mountain tops while others are restricted to the valleys. An island for one organism may not be suitable for another. The volcanic island, Surtsey also known as the island of fire, has made it possible for scientists to study the evolution of plants and animals naturally. It was named after the Norse mythological figure, Sutur, because he was said to carry a sword of fire. There is no more volcanic activity feeding this island, meaning the peak of the island is set at 508 ft. above sea level. Due to porous scoria (very porous volcanic rock); 3 acres of land disappear every year. When the island first came about Scientists seized their opportunity to see just how long it would take for life to form on this new and untouched land. Within the same year a searocket was found growing on the shoreline. These plants most likely came by sea from the island Heimaey, which is 11 miles away (Kuschk). Birds started colonizing on the island instead of just using it as a stop during migration, so vegetation started to flourish. Birds are thought to be 75% responsible for all seed dispersal. There is now grassland 25 acres long that can be seen from satellite due to gulls colonizing in a specific area. So far, 69 species of plants have been documented on Surtsey and 32 have become entrenched, meaning they’ll always be on the island (Kuschk). Melissa Gantt Bio 111 D6 Page 5 Insects have been on Surtsey since the beginning; starting with flying bugs, then spiders and so on. They came in contact with this new land mass via drift wood, dead carcasses that washed up; and by grass and soil that broke off from other land masses and floated to its shore. Not only are birds a main occupier, seals have been using the island since 1983 for breeding grounds. The colonization is still happening with chestnut worms and slugs, found in 1990, which helps fertilize the soil. The significance and evolutionary possibilities of this island led to a ban on all non-science related visitation. During mid-1970, someone improperly discarded their bowel movement and a tomato plant took root (Kuschk). Another incident occurred when boys from the neighboring island Heimaey planted potatoes. Both of these foreign plants were removed. This way nothing is growing there due to human interference. Melissa Gantt Bio 111 D6 Page 6 Surtsey is giving us a great look at evolution in the process. From the beginning in 1963, it was created from a volcanic erpution. Birds started landing on the island during migration and dropped off seeds. Eventually, a group of gulls colonized and there is now 25 acres covered in grass. This means birds, inclement weather, and ocean tides all made this new, untouched land habitable; naturally. There was very minimal human interference and this island is thriving with plants and wildlife. The lead scientist on the island, Borgthor Magnusson, was just 11 years old when he saw the volcano erupt. He stated, “this opportunity to follow the colonization of a new site on earth right from its formation – from the time when it was a pile of ash until now, with many hundreds, even thousands of plant species and animals, to be able to follow that is very, very rare” (Blask). There is, however, a possibility of seeing this phenomenon again. A volcanic eruption is currently causing a new island. They are uncertain if the eruption in the Pacific will last long enough to create a sustainable island. It is only 650 ft. wide and 980 long as of lately. Unfortunately, the opportunity to study evolution at this potential island is being overlooked by territory disputes between China and Japan. Melissa Gantt Bio 111 D6 Page 7 Mountains can be considered islands because they have a wide range of habitats for different species. The Rocky Mountains is a great example of this, because they range in western North America, from far north Canada down to southeastern New Mexico. This creates many temperatures and rainfall types. There is a mixture of habitats in the Rockies. They consist of alpine, subalpine, and boreal throughout the Northern Rockies (Ecology). Coniferous forests are located throughout the range which, “contain the highest level of biomass and are notable for their trees” (Ecology). There are also wetlands and prairies where the Rockies meet the plains. This mountain range holds a huge variety of wildlife; so much so that ecologists divided it into life zones. They figure this out by seeing where trees are supported and species that define traits and characteristics of the environment, known as indicator species (Ecology). “Piñon pine and juniper are common in the southern Rockies, while they give way to firs, pines, and spruces further north. Beyond the timber line (the altitude above which trees cannot grow), mountain goats and bighorn sheep browse in alpine meadows. The forests downslope are home to bears, deer, elk, hares, minks, cougars, porcupines, squirrels, and other woodland creatures. The rivers of the region are rich with fish such as rainbow trout and grayling. Within the Rocky Mountains lies the Continental Divide, which separates the waters that flow to the Pacific Ocean from those which will flow into the Atlantic . . .” (Rocky). Just looking at the flora and fauna living in each zone provides Darwin’s theory with proof of colonization and isolation. Not only are there places where trees will or will not grow, but there are places where different animals are unable to live. Most likely the squirrel would have died off if they lived high in the alpines. However, they either would have adapted to their environment or migrated down to the valleys to find a more suitable one. Melissa Gantt Bio 111 D6 Page 8 Why does the bighorn sheep choose to live on the mountain tops? They have to have rough hooves just to get the right amount of traction to maneuver on the side of a mountain. Some study suggests bighorns could have possibly adapted to the mountain tops to avoid predators such as coyotes, but these are still just theories. We weren’t around when these species settled and fully adapted to their habitat. Now that we are established, permanent human settlement in the Rocky’s is forcing species to decline. Gray wolves and grizzly bears were completely eliminated from the United States and are only returning because of conservation measures in affect. Humans have become a main reason for a decline in what once was a thriving species. Because of these actions we are now considered a super predator. Human beings have forced evolution to occur because of a continuous population and just out of pure ignorance. “[The] Bighorn Sheep is one of the many species documented as getting smaller, on average” (Britt). This has forced body size and reproduction ability to occur 300 times faster than what’s naturally acceptable. One reason larger species are dying off is because of trophy hunters, people that kill to boast and keep horns as evidence. Basically animals are being killed for their traits and features. Once again, like with the Pinta Tortoise, people are exposing these species adaptation and killing them off. Unfortunately, it’s not for food this time around, but just for a status in the hunting community. This is why Surtsey Island is so important. Not only are we experiencing evolution first hand, but there is a ban on anyone setting foot on the island other than scientists to prevent unnecessary change. Surtsey is allowing us to study the evolutionary changes from the beginning, starting with nothing but a volcanic eruption. Melissa Gantt Bio 111 D6 Page 9 We now have a habitable environment consisting of birds, plants, seals and recently discovered slugs and earthworms. Everything that has occurred on this island is happening through nature. The seeds that washed up were quite large, capable of floating and can tolerate salty water, which is how they were able to make landfall and germinate. Woolly Willow, Dwarf Willow, and Tea-Level Willows are now all established on the island, because these species have tufts of hair making it easier to be dispersed long distances through wind (Ólafsson). Birds also played a major role in creating an adaptable environment. Originally the island was used just during migration. Once Scurvy grass appeared where the gulls rested it made more sense for these gulls to colonize there. Hopefully, Surtsey will continue to grow and with the right protection measures we will see natural selection occur on this island and get some real answer about evolution as we know it. “A total of 300 different species of invertebrates have been found on Surtsey, and approximately half of them have become permanently established” (Ólafsson). Flying insects are the most obvious of species. One of interest is the Ceutorrhyncus insularis, a very small snout beetle that feeds on scurvy grass and is an extremely rare species on a global scale. Melissa Gantt Bio 111 D6 Page 10 A look at Island Geography forming Melissa Gantt Bio 111 D6 Page 11 A very interesting island that has been mistaken for a continent before is Greenland. It is known for being the world’s largest island. One main reason it is considered an Island is because it sits on North America’s tectonic plate and is not geologically separated from Canada, the U.S., and Mexico. It is mostly covered by an ice sheet, making two-thirds of the land subject to permafrost. This island is very important when it comes to climate change. It will be able to raise sea levels dramatically if it were to melt via global warming. Interestingly enough all cities in Greenland are built on the coast. That is the only places where there is no ice formed. Greenland was first discovered by Erik the Red who was exiled from Iceland for 3 years for committing murder. He named this island Groenland, which translates into “green land.” There were rumors he named it this to attract more settlers to the area, thinking people would be willing to travel if the name sounded appealing. Being very persuasive he was able to get 25 fleet ships to follow him back to this “green land.” There were women, children, and domestic animals on board. Only 14 ships made it to their destination. They then separated into 2 colonies, known as the east and west settlements. This icy land actually restricts all human activity to the coasts. 25000 BC is thought to be the year humans first set foot on the island. However, when the Icelandic Vikings settled on the land it looked to be uninhabited. They took settlement along the southwestern coast. “Excavations have shown that the fjords at that time were surrounded by forests of 4-to 6-metre-tall birch trees and by hills covered with grass and willow brush” (History of). However, it is thought that when the Norse landed they cleared out the trees for fuel and building material, and allowed their goats to graze during winter and summer, clearing out much of the grass originally there. Melissa Gantt Bio 111 D6 Page 12 During the 14th and 15th centuries the “Little Ice Age” began. The Inuit, better known as Eskimos in the U.S., survived icy weather and the “Little Ice Age” period. However, the Norse settlements were disappearing so fast that Denmark-Norway still claimed sovereignty on the land (History of). The climate survival was very difficult. However, this led to one culture in particular being able to succeed as other groups died out. The Inuit were the only people inhabiting Greenland for several centuries. The ice-free areas benefited a few cold tolerant species to survive the Ice Age. Other than that, it can be assumed that all life was exterminated and current species invaded Greenland after the Ice Age ended. As of now Greenland’s population is 89% Inuit and 11% Danish (Upton). The majority of Greenland’s plant species originated from Europe. However, their national flower, Niviarsiaq, meaning “young lady”, is commonly found in North America. The fauna such as reindeer, polar bears, and artic fox can be found living on Greenland and other land. There are not many unique animal species to the island due to the extermination of animals during the Ice Age. There are 10 unique vascular plants found nowhere else but in Greenland, including East Greenland Saxifrage Greenland Alkali Grass Porsild’s Alkali Grass Greenland Fescue Greenland Blue-eyed Grass Greenland Pondweed Two Pussy Toes 15 endemic micro species of hawkweed. Melissa Gantt Bio 111 D6 Page 13 “The evolution of vascular tissue allowed for an early dominance of these plants on land. It gives them the ability to transport water and dissolved minerals through specialized strands of elongated cells that run from the plant root to the tips of the leaves” (Vascular). Just like the first plants that came on land. There is also a flatworm, and a comb jelly from Umanak Fjord that is native to Greenland. These are two marine invertebrates considered to represent families known only from Greenland (Greenland). Adaptation to cold and drought are main factors in the distribution of species. The Greenland shark is a one of a kind animal that can be found throughout the northern Pacific Ocean at great depths. It is one of the only fish that prefer artic to subarctic temperature ranging from -2 and 7 degrees Celsius at all times. During winter they can be found around the surface and near the edge of ice flows. Their skin color can range from brown to a violet. These sharks can be up to 21 feet in length, but have ironically small fins. There dorsal fin is equal in size, sits on the middle of the sharks trunk and is spineless (Rogers). Melissa Gantt Bio 111 D6 Page 14 Another interesting fact about these historic sharks is their eyes. They have spiracles on the tops and bottoms which allow them to breathe when buried in sand or laying on the bottom of the floor. However, most of these sharks are blind due to parasites living off their eyes. They use to be hunted down for their liver in Greenland, Norway, and Iceland, but that is no longer practiced. Inuit hunters do eat the shark but it is not a delicacy due to the cold water they live in and the fact the meat is poisonous when raw. Unlike the Bighorn Sheep and the Pinta Tortoise, hunters are not putting these sharks into extinction. Unfortunately many other shark species aren’t as lucky. Conclusion Charles Darwin once said, "in the struggle for survival, the fittest win out at the expense of their rivals because they succeed in adapting themselves best to their environment." Reflecting back on my research of Island Geography I saw this quote being fulfilled time and time again. Just look at the Galapagos Islands, the initial arrivals served only one purpose; to die so they could feed the flora for the next generation. Slowly but surely this Island came to have over 25 new species of finches along with Pinta Tortoises that were able to adapt after the rat infestation. If not for humans who knows how the Pinta Tortoise could have evolved. Even now Surtsey Island is showing us evolution from the beginning, starting with a pile of ash to becoming colonized by gulls who helped scurvy grass grow over 25 acres of land. This will give us a look at natural selection, with very minimal human interference. Birds, weather, ocean tides, natural selection, and even mutation can form and create colonization on an island. Looking at the huge variety of species and geography of these islands shows why Darwin was so intrigued by the Galapagos and its finches. Melissa Gantt Bio 111 D6 Page 15 Island Geography can be found all over the world from the peaks and valleys of mountains to the depths of the Pacific Ocean. Species are constantly adapting and evolving to geographical and ecological changes. The same species of finches might have adapted different features if they were located on Greenland or could have died off to make way for the fittest of the species. Natural Selection and Evolution are able to be noted by looking at the various “islands” I mentioned throughout this paper. Darwin was lucky enough to stumble across many types of species of finches that adapted to best benefit them. We are once again able to see these evolutionary traits in motion all thanks to Surtsey Island. Eventually we will start seeing positive mutations in DNA and hundreds of thousands of flora and fauna will continue to flourish. Melissa Gantt Bio 111 D6 Page 16 Works Cited “Biological Evolution of the Galapagos.” Galapagosislands.com. Web. 25 Nov 2013. Blask, Sara. “Iceland’s New Island is an Exclusive Club—for Scientists Only.”Csmonitor.com. 24 Oct 2008. Web. 28 Nov 2013. Britt, Robert Roy. “Super-Predators: Humans Force Rapid Evolution of Animals.” Livescience.com. 12 Jan 2009. Web 2 Dec 2013. “Buffon, Georges-Louis Leclerc, Comte De." Complete Dictionary of Scientific Biography. 2008. Encyclopedia.com. 26 Nov. 2013. “Darwin’s Finches.” Avibushistoriae.com. Wikipedia. Web. 25 Nov 2013. “Ecology of the Rocky Mountains.” En.wikipedia.org. Wikipedia. Web. 1 Dec 2013. “Greenland.” Intreasures.com. Living National Treasures. Web. 4 Dec 2013. “History and Achievements.” Darwinfoundation.org. Charles Darwin Foundation. Web. 27 Nov 2013. “History of Greenland.” En.wikipedia.org. Wikipedia. Web 5 Dec 2013. “Island Biogeography.” En.wikipedia.org. Wikipedia. Web. 28 Nov 2013. Kuschk. “Surtsey: Evolution of an Island.” Basementgeographer.com. 28 May 2012. Web. 1 Dec 2013. Ólafsson, Erling. “Colonization on the Island.” Surtsey.is. Surtsey: The Surtsey Research Island. 6 May 2007. Web. 3 Dec 2013. “Overview.” Islandbiogeography.org. National Science Foundation. Web. 26 Nov 2013. “Rocky Mountains.” World-geography.org. World-Geography. 7 July 2010. Web. 1 Dec 2013 Melissa Gantt Bio 111 D6 Page 17 Rogers, Mike. “The Greenland Shark.” Sharksider.com. Shark Sider. Web. 5 Dec 2013. Upton, Emily. “Why Greenland is an Island and Australia is a Continent.” TodayIfoundout.com. Today I Found Out: Feed your Brain. 15 April 2013. Web. 5 Dec 2013 “Vascular Plants.” Newworldencyclopedia.org. New World Encyclopedia. Web. 4 Dec 2013. Watson; Oxford, Pete, and Graham. “Human Discovery: Charles Darwin.” Galapagos.org. 2009. Print. Web. 24 Nov 2013. Melissa Gantt Bio 111 D6 Page 18