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Video ■ Crash Course: Simple Animals – https://www.youtube.com/watch?v=tIfsHPpkSPs INVERTEBRATE EVOLUTION Essential Question: what are the major trends in invertebrate evolution? The first multicellular animals ■ Invertebrate fossils date back between 575 and 543 millions of years ago ■ Distinctive flat, plate-shaped ■ Showed little cell specialization or cephalization ■ Lived in shallow seas and some may have had photosynthetic algae living within them Beginnings of invertebrate diversity ■ A few million years later (Cambrian Period) radically different invertebrates ■ Some animals developed shells, skeletons, and other hard body parts, providing a wealth of fossils ■ Ancestors of most modern animal phyla first appeared Feeding and digesting ■ Simpler animals break down food primarily through intracellular digestion (food digested inside the cells) – Ex. Sponges ■ Complex animals use extracellular digestion (food broken down outside of cells and then absorbed into the body) – Ex. Mollusks, annelids, arthropods, and echinoderms ■ Flatworms and cnidarians use both Feeding and digesting ■ Cnidarians and most flatworms ingest and expel waste through the same opening ■ Complex animals digest food in a digestive tract and expels waste through the anus ■ Specialization of the digestive tract allows food to be process more efficiently Respiration ■ Respiratory organs have large surface areas that are in contact with the air or water. ■ For diffusion to occur, the respiratory surface must be moist ■ Many simple aquatic animals respire through their skin. Respiration ■ Complex aquatic animals develop gills (rich in blood vessels that brings blood close to the surface for gas exchange) ■ Terrestrial animals have respiratory surfaces covered in either water or mucus as well as having the air moistened as it travels through the body INVERTEBRATE EVOLUTION II Essential Question: How do different invertebrate phyla carry out life functions? Circulation ■ Most complex animals move blood through their bodies using one or more hearts ■ Open circulatory systems- blood is only partially contained within a system of blood vessels. One or more hearts pump blood through blood vessels into a system of sinuses or sponge cavities – Ex. arthropods and most mollusks Circulation ■ Closed circulatory systems- Heart or heart-like organs forces blood through vessels that extend throughout the body. Blood is kept at a high pressure and is more efficient than open circulatory systems – Ex. Annelids and some mollusks Excretion ■ Aquatic invertebrates diffuses ammonia from their body tissues into the surrounding water ■ Terrestrial invertebrates must conserve water so they convert ammonia into a compound called urea which is eliminated through urine Response (Nervous system) ■ Nerve net- simplest nervous system found in cnidarians that consists of individual nerve cells in a netlike arrangement throughout the body ■ Centralized- in flatworms and roundworms, the nerve cells are more centralized in small clumps in the head ■ Cephalization- in cephalopod mollusks and arthropods, the nerve cells (ganglia) are organized into a brain that controls the nervous system Response (Nervous system) ■ Complex animals may have a variety of specialized sensory organs that can detect light, sound, chemicals, movement, and even electricity – Ex. Flatworms have simple eyespots that detects the presence of light while arthropods have eyes that can detect motion and color, and can forms images Movement and support ■ Hydrostatic skeletons- muscles surround a fluid-filled body cavity that supports the muscles – Ex. Annelids and certain cnidarians ■ Exoskeletons- an external skeleton made of chitin in which muscles are attached inside the hard body – Ex. Arthropods and shelled mollusks ■ Endoskeletons- a structural support inside the body – Ex. Echinoderms Reproduction ■ Most invertebrates reproduce sexually, however depending on favorable environmental factors, many may also reproduce asexually (faster than sexually) ■ Hermaphrodites- individuals that produce both sperm and eggs – Ex. Earthworms