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Chapter 19 Sect 3
Biogenesis
Production of new or
living organisms. Living
things come from living
things.
Ex: Spider eggs become
spiders
Scientists Louis Pasteur
came up with this idea.
Endosymbiosis
Explains how eukaryotes
came from prokaryotes.
Primordial Soup
Theory
a solution rich in organic
compounds in the
primitive oceans of the
earth, from which life is
hypothesized to have
originated.
Miller Urey
Experiment
Used Earth’s early
atmosphere composition
to show that life can
spontaneously happen.
The Mysteries of Life’s Origins

What do scientists hypothesize about early Earth and the origin of life?

Geological and astronomical evidence suggests Earth forms as pieces of
cosmic debris collided with one another.

For millions of years, violent volcanic activity shook Earth’s crust along
with comets and asteroids bombarding the surface.

About 4.2 billion years ago Earth cooled allowing solid rocks to form and
water to condense and fall as rain.

Earth’s early atmosphere contained little or no oxygen. It was
principally composed of carbon dioxide, water vapor, and nitrogen, with
smaller amounts of carbon monoxide, hydrogen sulfide, and hydrogen
cyanide.

A few deep breaths would have killed you!
The First Organic Molecules 1/2

Could organic molecules assemble under conditions on early Earth?

Stanley Miller and Harold Urey tried to answer that question.

They filled a sterile flask with water and boiled it – illustrating the
oceans.

To the water vapor they added methane, ammonia, and hydrogen –
illustrating the atmosphere.

They passed the gases listed above through electrodes to simulate
lightning.

They then passed the gases through a condensation chamber where cold
water cooled them, causing drops to form. T

The liquid circulated for a week.

The end result was 21 amino acids!
The First Organic Molecules 2/2

Miller and Urey’s experiment suggested how mixtures of the organic
compounds necessary for life could have arisen from simpler compounds
on a primitive Earth.

Were they wrong???

Yes they were wrong. More research showed that the composition of
Earth’s early atmosphere was different than what they used in their
experiment.

But recent experiments using the correct ingredients in the atmosphere
still produced organic compounds.

In fact, in 1995 one of Miller’s more accurate mixtures produced
cytosine and uracil which are two bases found in RNA.
Formation of Microspheres

A stew of organic molecules is a long way from a living cell.

Geological evidence suggests that during Earth’s Archean Eon (200-300
million years ago) after Earth cooled enough to carry liquid water, cells
similar to bacteria were common. Where did these come from though?

Large organic molecules form tiny bubbles called proteinoid
microspheres under certain conditions.

Microspheres are not cells, but have characteristics of living systems.

They have selectively permeable membranes and have simple means of
storing and releasing energy.

Several hypotheses suggest that structures similar to proteinoid
microspheres acquired the characteristics of living cells as early as 3.8
billion years ago.
Evolution of RNA and DNA

Where did RNA and DNA come from?

Scientists haven’t solved this yet, but that doesn’t mean they won’t.

A number of experiments however simulating conditions on early Earth
show small sequences of RNA forming from simple molecules.

Is this good news? The answer is yes!

Scientists know that under the right conditions some RNA sequences
help DNA replicate, some RNA process mRNA, some catalyze reactions,
and some even grow and replicate on their own!

The “RNA World” hypothesis proposes that RNA existed by itself before
DNA. From this simple RNA-based system, several steps could have led
to DNA-directed protein synthesis.

It is still being tested today.
Production of Free Oxygen

Microscopic fossils of prokaryotes that resemble bacteria have been found in
Archaeon rocks more than 3.5 billion years old. These evolved in the absence of
oxygen because early Earth hardly had any oxygen.

During the Proterozoic Eon, photosynthetic bacteria became common. 2.2 billion
years ago, these organisms were churning out oxygen.

The oxygen combined with the iron in the oceans creating iron oxide (rust) and sank
to the bottom of the ocean forming great bands or iron that are the source of most
iron mined today.

Without iron, the oceans changed color from brown to blue-green.

Oxygen began to accumulate in the atmosphere, the ozone layer formed, and the
skies turned to blue.

Over several hundred million years, oxygen concentration levels rose to those of
today.

Early life forms died because they couldn’t breathe oxygen. Other organisms
evolved to breathe oxygen.
Origin of Eukaryotic Cells

What theory explains the origin of eukaryotic cells?

One of the most important events in the history of life was the evolution of
eukaryotic cells from prokaryotic cells.

How did these complex organisms evolve?

Researchers hypothesize that about 2 billion years ago some ancient prokaryotes
began evolving internal cell membranes.

Endosymbiotic Theory – prokaryotic cells entered ancestral prokaryotes. These
intruders didn’t infect their host and the host didn’t digest them.

Small prokaryotes began living inside larger cells.

The Endosymbiotic Theory proposes that a symbiotic relationship evolved over time,
between primitive eukaryotic cells and the prokaryotic cells within them.

This was proposed about a century ago. Looking under the microscope the
membranes of mitochondria and chloroplasts look like the cell membranes of free
living prokaryotes.
Endosymbiotic Theory

This was proposed about a century ago. Looking under the microscope
the membranes of mitochondria and chloroplasts look like the cell
membranes of free living prokaryotes.

This led to two hypotheses.

The first says that mitochondria evolved from endosymbiotic
prokaryotes that were able to use oxygen to generate ATP. These energy
generating prokaryotes evolved into mitochondria.

The other hypothesis says that chloroplasts evolved from endosymbiotic
prokaryotes that had the ability to do photosynthesis.

Over time these prokaryotes turned into chloroplasts of plants and
algae.
Modern Evidence
 During
the 1960s, Lynn Margulis gathered evidence supporting
the endosymbiotic theory.
 Margulis
found that mitochondria and chloroplasts contain DNA
similar to bacterial DNA.
 She
also found that mitochondria and chloroplasts have
ribosomes whose size and structure look similar to that
bacteria mentioned above.
 Mitochondria
and chloroplasts reproduce through binary fission
when cells containing them divide by mitosis.
 These
similarities show strong evidence.
Sexual Reproduction and Multicellularity

What is the evolutionary significance of sexual reproduction?

Sometime after eukaryotic cells arose, they began to reproduce sexually.

The development of sexual reproduction sped up evolutionary change because
sexual reproduction increases genetic variation.

Asexual reproduction produces identical genetic cells.

Sexual reproduction leads to a “shuffling” of genes which is why offspring look
different from their parents.

Genetic variation increases the likelihood of a population’s adapting to new or
changing environmental conditions.

Multicellular organisms evolved a few hundred million years after the evolution of
sexual reproduction.

Early multicellular organisms underwent a series of adaptive radiations, resulting in
greater diversity.