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1 Organisms and Energy Energy and Nutrients All living things need energy Energy is the capacity to do work or to cause change chemical energy: energy stored in the structure of molecules organisms can use some of this energy as its released during chemical reactions, but most of it is released as unusable energy free energy: energy that’s available to do work examples of free energy- energy plants use for growing and producing food and the energy you use for exercise and thinking living cells need a constant source of free energy for chemical, mechanical, and transport work transport work: involves the movement and concentration of the nutrients needed to make complex molecules and to increase cellular organization during growth mechanical work involves movement like muscle contractions that enable you to kick a ball chemical work involves maintenance of the cell – cells need to manufacture molecules to replace ones that are used up or damaged nutrients: a substance that supports the growth and maintenance of an organism ex: carbs, lipids, proteins heterotrophs: an organism that obtains its food molecules (carbon compounds) from other organisms consumers: an organism that feeds on other organisms or on their organic wastes **heterotrophs = consumers** ex: animals, fungi (mushrooms and mold), and most bacteria obtain the nutrient ions from decomposers by eating plants decomposers: any organism that lives on decaying organic material, from which it obtains energy and nutrients ex: bacteria, fungi, mineral ions like potassium, iron, nitrate, and ammonium complete the breakdown of organic nutrients and return inorganic nutrients to the soil or water, where they are available for reuse autotrophs: an organism that forms its own food molecules (carbon compounds) from abiotic materials producers: any organism that produces its own food **heterotrophs = consumers** ex: plants, protists, some bacteria they absorb the nutrient ions dissolved in soil water from decomposers through their roots photoautotrophs: use the process of photosynthesis to make their own food chemoautotrophs use the process of chemosynthesis to capture energy, which is stored as chemical energy and used for cell work ©SarahStudyGuides 2 photosynthesis: the process where cells use light energy from the sun to make organic compounds from inorganic compounds sunlight CO2 + H2O chlorophyll C6H12O6 + O2 Chemical energy chlorophyll- allows plants to capture solar energy and change it into chemical energy photosynthesis converts light energy into chemical energy and creates glucose and oxygen chemosynthesis: a biological pathway that uses energy from the oxidation of inorganic substances to drive the formation of organic molecules cellular respiration: the series of chemical reactions by which a living cell breaks down carbohydrates and obtains energy from them cellular respiration changes chemical energy produced by photosynthesis into free energy in the form of ATP for the cell to use for cell work it produces carbon dioxide and water as waste products cells: 90% heat energy C6H12O6 + O2 enzymes Chemical energy CO2 + H2O + energy nd 2 law 10% free energy 1st law 1st law of thermodynamics: 1) 2) 3) 4) 5) 6) reproduce movement move materials in and out of cells chemical reactions growth maintenance Energy can’t be created or destroyed, but it can change forms 2nd law of thermodynamics: Most of the energy available is converted into unusable energy in the form of heat energy Systems tend to change in a way that increases the disorder, or entropy, of the system plus its surroundings Sunlight is the source of all energy because only autotrophs can capture energy from inorganic sources, autotrophs directly or indirectly supply the energy and organic nutrients needed for all heterotrophs heterotrophs must eat to get energy and nutrients to grow and develop sunlight producers primary consumers heat secondary consumers heat tertiary consumers heat quaternary consumers heat this chain can’t continue endlessly because it would run eventually run out of energy *the 2nd law of thermodynamics* every time energy travels from level to level, 90% of it is lost as heat: only 10% of the energy transferred to each level makes it the amount of energy gets smaller as levels increase and the world gets more disordered Energy and nutrients flow from the producers to consumers to decomposers tertiary consumers abiotic factors: nonliving things in an ecosystem; referring to aoverlapping physical orfood nonliving component of an Food web: the chains of an ecosystem secondary consumers ecosystem Food chain: each individual track of energy transfer primary consumers ©SarahStudyGuides Ex of a food chain: producers grass mouse snake hawk eagle quaternary consumers 3 ex: soil, minerals, water, weather, climate, biotic factors: living things in an ecosystem; referring to a living component of an ecosystem ex: the organisms in an ecosystem ecosystem: a community of living things and its abiotic environment made up by its abiotic and biotic factors has lots of factors in common ex: ocean, marshes, coral reefs, desert, rainforest, pond, prairie, forest habitats: type of place where particular organisms live ex: in a pond- some organisms are bottom-dwellers and others live along the shore (these are different habitats) biosphere: the outer portion- air, water, and soil- where life is found ex: our earth all ecosystems combined to make up earth’s biosphere Energy and Entropy entropy: how much energy in a system has become so dispersed that it’s no longer available to do work organisms must be well organized to remain alive and to grow a key to maintaining organization of all living systems is energy in ecosystems, light or chemical energy flows from the environment (from the sun or inorganic compounds) to producers to consumers to decomposers producers don’t use all the energy they absorb in photosynthesis- most is released as heat energy same with consumers- some of the chemical energy is converted into free energy for cell work, but most is released as heat energy as energy flows through food webs, it escapes into the surroundings in the form of heat energy so there’s only a one-way flow of energy through food webs living systems overcome the tendency toward entropy by constantly obtaining energy from their surroundings organisms stay organized and can function and grow only as the entropy of their surrounding increase so: energy isn’t created or destroyed so the total energy of the universe remains the same (1st law) and it’s dispersed as heat energy which increases the entropy of the universe (2nd law) Enzymes and Energy To release chemical energy to perform work, cells must have a way to break and form chemical bonds Chemical reactions in most organisms take place within a narrow range of temperatures Those temperatures aren’t high enough to supply the activation energy to start a reaction Many organic compounds are sources of energy for the cells of living organisms – 3 most important energy sources are monosaccharides, disaccharides, and polysaccharides The first step in releasing chemical energy from organic compounds is to break them down into their building blocks: glucose, amino acids, fatty acids and glycerol ©SarahStudyGuides 4 This chemical energy released from organic compounds is obtained by breaking chemical and then forming new chemical bonds One indication that organisms release and use energy is that all organisms give off heat. Some like fireflies, also give off light Energy will be released in a chemical reaction if the chemical bonds that are broken hold more energy that the new ones that are formed What are Enzymes? enzymes: proteins- tertiary level globular or spherical they speed up chemical reactions by lowering activation energy activation energy: the amount of energy needed to start a reaction any chemical reaction requires the input of energy called activation energy enzymes are catalysts catalysts: chemicals that lower activation energies chemical reactions would otherwise be too slow to sustain life mixing reactants won’t produce a product – the only way they can be changed is by a biochemical reaction, again requiring the action of an enzyme each enzyme only catalyzes specific reactions the specific reaction catalyzed by an enzyme depends on a small area of its tertiary structure called the enzyme’s active site active site: the indent or groove where the substrate binds with the enzyme the active site’s shape must match the shape of the substrate substrate: the starting molecules the close fit of the active site and substrate brings the enzyme and substrate close together which lowers the activation energy with allows the chemical reaction from the substrate to the product to happen the fit of the enzyme and substrate is called the enzyme-substrate complex denatured: doesn’t work anymore because the shape of the active site is changed because the shape of the active site is changed, it doesn’t match the shape of the substrate anymore, so the enzyme and substrate can’t combine to form an enzyme-substrate complex two things denature enzymes: 1. temperature (like in the seed lab) 2. pH (like in the digestive system) enzymes aren’t rigid so sometimes the shape of the substrate causes it to adjust its shape slightly for a better fit between the enzyme and substrate the reactions don’t consume the enzymes, so they are reusable and remain unchanged once the chemical reaction occurs, the new product molecules break away, leaving the enzyme the same as it was before the reaction so they can be reused these enzyme-catalyzed reactions are reversible-- it can build substrates into a product or break a product into substrates enzymes can speed up chemical reactions in both directions ©SarahStudyGuides 5 How do Enzymes affect Chemical Reactions? 1) speeds up reactions 2) allows reactions to occur at lower temperatures ex: we burn glucose, but we aren’t 350º 3) can build up or break down substrates 4) enzymes have specificity – each substrate has a specific enzyme ex: the enzyme in saliva is amylase and it breaks down the substrate amylose 5) enzymes must fit the substrate at the active site 6) enzymes’ active site can be altered by changes in pH and temperature which means the enzyme becomes denatured denatured: doesn’t work anymore 7) have an “-ase” ending Chemical Reactions in Organisms chemical reactions occur continuously in all organisms metabolism: consists of all the chemical activities and changes that take place in a cell or organism there are two types of metabolism (and they all involve enzymes): synthesis – “building up” reactions decomposition – “breaking down” reactions synthesis reactions: include biosynthesis reactions that form larger, more complex molecules from small, less complex ones ex: glucose forming starch, nucleotides forming DNA, amino acids forming proteins proteins building tissues, like blood and muscle photosynthesis building sugars from carbon dioxide and water biosynthesis requires free energy, because the products are more ordered and contain more chemical energy than the simpler, less ordered reactants since the reactants added together make the product and the reactants have less energy than the products, free energy needs to be added to the reactants to complete the product biosynthesis also enables organisms to grow and maintain their structure decomposition reactions: large molecules break down into smaller molecules ex: glycogen breaking into glucose in muscle cell glucose breaking down into carbon dioxide and water during cellular respiration the energy stored in the glucose becomes available to the muscle cell or for other biosynthesis reactions decomposition releases free energy and some heat energy in the form of ATP and used by the cell because they produce simple molecules from complex molecules, which increases entropy cells use some of the free energy and simple molecules released during decomposition for biosynthesis of other macromolecules ©SarahStudyGuides 6 so synthesis and decomposition reactions in the cell are coupled to energy flow and the cycling of matter Energy Transfer and ATP Through a process of decomposition called oxidation, certain bonds are broken and rearranged oxidation: the removal of electrons from a molecule some of the energy of the original molecule is released as heat energy and free energy the free energy follows a series of electron transfers and ends up as ATP What is ATP? Adenosine triphosphate ATP stores energy that can be transferred to other molecules drives the reactions of biosynthesis connects many energy-conversion reactions during metabolism has been called the “energy currency” of living cells just like people have to pay a fee to exchange foreign currency to our currency: cells exchange chemical energy for ATP—the “fee” is the energy lost as heat energy ATP than pays most of the energy “debts” inside a cell ATP is a nucleotide consisting of 3 parts: adenine, ribose, and 3 phosphate groups the structure of ATP makes it an efficient and useful energy-transfer molecule in cells ATP supplies much of the energy for the transport and mechanical work needed by cells when an ATP is involved in a chemical reaction, the bond between the 2nd and 3rd phosphate groups break and free energy is released high energy! O- O- OO – P –– P –– P –– O O O *the oxygens repel each other lots energy are in the bonds to hold them when these bonds are broken, lots of energy explodes *single covalent bonds *when this bond is broken (the 2nd bond), lots of energy explodes ATP is continually synthesized and broken down in cells ATP ADP + P + energy (decomposition) a high amount of energy is released when the bond holding the last phosphate is broken this is the reason that ATP makes energy this energy is free energy and provides energy for cell work if the bond between the 1st two phosphate bonds is broken, less energy is released than breaking the bond between the 2nd and 3rd phosphate groups ADP + P + energy ATP (synthesis) To form ATP again, ADP must be combined with one phosphate group which requires free energy ©SarahStudyGuides 7 produced (synthesized) from glucose decomposed into ADP (adenosine diphosphate) 90% heat energy Glucose enzyme synthesis glucose-6-phosphate + O2 energy ATP decomposition ADP CO2 + H2O + energy for cell cork 1. 2. 3. movement growth reproduction when a phosphate is transferred to glucose, glucose-6-phosphate is formed this molecule is easily converted by cells into energy to form glucose-6-phosphate, the synthesis reaction of glucose must be coupled with decomposition reaction of ATP for energy ATP was decomposed to form ADP + P + energy At the same time, glucose was being synthesized Before glucose can be completely synthesized, it has to accept the phosphate group and free energy from the decomposition ATP The phosphate group from the decomposition of ATP attaches to the 6th carbon in glucose and the free energy activates the reaction the product is glucose-6-phosphate It’s called glucose-6-phosphate because the phosphate group from the decomposition of ATP attaches to the 6th carbon in glucose Cells can use the free energy stored in ATP to supply activation energy or start reactions 1 glucose molecule can be converted into approximately 36-38 ATP molecules you will eventually run out of ATP if you don’t eat or have oxygen ex: when you run and run, you’ll eventually have to stop due to lactic acid which makes your muscles burn and the fact that you need oxygen for ATP Digestion Digestion Inside and Outside Cells only small molecules can pass through membranes and into cells, where energy release takes place however, most food particles that animals ingest are too large so they must break them down into macromolecules like carbs, proteins, and lipids, and then break them down even further digestion: the processes that break down food the smallest products of digestion: o carbs: glucose o lipids: fatty acids and glycerol o proteins: amino acids physical digestion: the breakdown of large pieces of food into smaller ones ex: chewing and grinding ©SarahStudyGuides 8 increases the surface area of food, making the chemical digestion part easier because the large surface area allows enzymes more access to the food particles chemical digestion: involves the breakdown of complex food molecules into simpler ones ex: starch becoming glucose enzymes in the various organs of the digestive tract control these chemical reactions without chemical digestion, nutrients obtained from large food molecules couldn’t be absorbed and used by the organism extracellular digestion: digestion that takes place outside of cells most animals, including humans, rely on this type of digestion most animals secrete digestive enzymes into a digestive cavity, where chemical digestion yields the simpler molecules that are then absorbed by the cells intracellular digestion: the digestion that takes place inside of cells with the foods the organisms made themselves plants digest food this way digestive enzymes break down the food into small molecules that the cell can use some plants have the ability to capture insects and digest them in special cavities formed by the leaves after the insect is trapped, the leaf cells secrete enzymes that digest it many organisms produce enzymes that digest food outside the organism itself and then absorb the nutrients into the cells ex: most fungi digest materials from dead plants and animals, like bread mold bread mold is a fungus that secretes enzymes that diffuse out of the cells and digest bread. The mold then absorbs the products of digestion into its cells. amino acids, sugars, minerals, water, and oxygen can diffuse into cells, but large molecules can’t diffuse into cells and are absorbed by other means complex multicellular animals digest food in specialized cavities or digestive tubes with two openings food enters the mouth at one end of the tube, and material that can’t be digested passes out of the anus at the other end of the tube the result is one-way movement of food and waste ex: earthworms cellulose from plants, which makes up grass, is so difficult to digest because cellulose is made of beta linkages, which most animals can’t digest because they don’t have the enzymes to digest it however, there are special adaptations that allow animals to digest cellulose -goats and cows have 4-chambered stomachs -horses and rabbit’s stomachs have special side pockets where microorganisms live and help digest cellulose carnivores can’t digest cellulose and consume mostly meat, which is more easily digested than grass, so they have shorter digestive tracts ex: lions have shorter digestive tracts than zebras because lions are carnivores and eat meat and zebras eat grass which is made of cellulose An Overview of Human Digestion ©SarahStudyGuides 9 ingestion: the process of taking food into the digestive tract ingestion starts in the oral cavity, or mouth is mostly water pH is almost neutral (6.8-7.2) two structures in the mouth physically digest food: - teeth: chewing action which increases the surface area able to be broken down faster surface area does this because… for example: imagine how long it would take to burn a piece of paper if you lit the corner now imagine if you tore up the paper into many pieces and lit each piece’s corner -the highly muscular tongue: allows food to come in contact with the saliva; moves food around carbohydrate digestion begins here: salivary glands: a watery secretion containing digestive enzymes that begins chemical digestion saliva: water + amylase salivary amylase: an enzyme that digests the substrate amylase (starch) to shorter polysaccharides and maltose (*starch isn’t fully broken down yet – it needs to be broken into glucose!) this is why when you keep food like a cracker in your mouth for a long time, it starts to taste sweet – the starch begins to break down into sugars when you swallow food, it passes over the epiglottis epiglottis: a trapdoor-like tissue that normally prevents food and liquids from entering the larynx or trachea (or airway) food then enters the esophagus esophagus: a muscular tube connecting the mouth to the stomach made of smooth muscle carries food to the stomach through peristalsis peristalsis: wavelike contractions that move food from the mouth to the stomach food then passes through the cardiac sphincter cardiac sphincter: the structure that regulates the movement of food from the esophagus into the stomach; its a ring-like muscle that acts like a valve relaxes, releasing small amounts of partially digested food into the stomach the food then reaches the stomach made of smooth muscle pH is really low- 2-3 denatures the salivary amylase (it no longer fits the substrate) infolding or ridges in the stomach called rugi -increases surface tension more gastric glands -leaves room for expansion ©SarahStudyGuides 10 digestion of proteins: the enzymes that break down large protein molecules in the stomach require a strongly acidic environment this acidic condition is provided by the stomach glands that secrete HCl the acid is so concentrated that it could destroy living tissue the cells of the stomach lining aren’t harmed because some of them secrete a thick, protective coat of mucus scientists now know that ulcers are caused by bacterium, not acid, and can be treated with antibiotics food stays in the stomach—presence of food secretes gastrin which secretes HCl which secretes pepsin Come gastrin: secretes HCl from HCl: makes the stomach really acidic which means carbs can’t be digested here; secretes pepsin gastric pepsin: breaks peptide bonds – takes polypeptide chains and breaks then into amino acids glands contractions of muscles lining the stomach wall thoroughly break up and mix food with secretions from stomach glands these secretions called gastric juices are composed of enzymes, mucus, and acid as a result of their action, the contents of the stomach soon become souplike after an average meal, the stomach usually empties in approximately 4 hours food then passes through the pyloric sphincter pyloric sphincter: the structure that allows food out of the stomach and into the small intestine like toothpaste coming out Accessory organs provide digestive material to break down substances even though they aren’t part of the digestive tract are organs that aren’t a part of the digestive tract but are important in secreting chemicals that help to digest food and contribute digestive juices to the small intestine through ducts: -liver (secretions enter gallbladder) lipids: bile – emulsifies fats – large fat droplets broken down into smaller fat droplets (*lipids aren’t completely digested yet!) -gallbladder (secretions enter small intestine) - pancreas (secretions enter small intestine) pH of 7-8 carb, lipid, and protein digestion are all completed here! carbs: amylase – breaks down amylose into glucose lipids: lipase – breaks lipids into 1 glycerol and 3 fatty acids proteins: trypsin – breaks down small polypeptides into amino acids food then enters the small intestine ©SarahStudyGuides 11 small intestine: a tube approximately 6 m long where chemical digestion is completed and food molecules are absorbed pH of 7-8 1st third involved in chemical digestion of carbs, lipids, and proteins 2nd third is involved in absorption of nutrients villi- dramatically increase the surface area to absorb nutrients food molecules are absorbed through the intestinal walls into the bloodstream the blood carries the molecules to all the cells where they are used in metabolism any undigested material eventually passes to the large intestine, where bacteria help produce several vitamins, gases, and other compounds reabsorbs water collects undigested wastes the vitamins and much of the water that was mixed with the food are absorbed through the walls of the large intestine these absorption partly dries out the wastes, called feces the feces are then eliminated through the anus feces: poop fiber, found in fruits and vegetables, helps with the elimination of wastes from the large intestine Carbohydrates, Proteins, Fats, and Absorption Carbohydrate Digestion begins in the mouth with the action of salivary amylase and is completed in the small intestine salivary amylase: an enzyme that digests the substrate amylase (starch) to shorter polysaccharides and maltose by breaking the chemical bonds in starch molecules and adding water molecules to the products of this breakdown salivary amylase amylose + H2O maltose saliva has a pH between 6-7.4, so salivary amylase functions best in that pH range but the stomach has a very low pH of 2-3 so carbohydrates can’t be digested in the stomach carbohydrate digestion is completed in the small intestine with pancreatic amylase the pancreas delivers pancreatic juices that convert the acidic food mixture to a basic pH again the maltose produced by salivary amylase is further broken down into glucoses by pancreatic amylase final result: glucose – done with carb digestion! Protein Digestion occurs in the stomach and in the small intestine first in the stomach: the presence of food causes cells to release a hormone called gastrin which enters the bloodstream ©SarahStudyGuides 12 gastrin secretes HCl which secretes pepsin in an inactive form called pepsinogen HCl changes pepsinogen to active pepsin pepsin: the active protein digesting enzyme in the stomach; it breaks large protein molecules into smaller polypeptides then further digestion in the small intestine: breaks these smaller polypeptides into amino acids trypsin is the intestinal enzyme that completes protein digestion by breaking peptide bonds, producing amino acids from those smaller polypeptides a basic pH activates trypsin and other intestinal chemicals a basic pH is necessary for the intestinal enzymes to function when food enters the small intestine, pancreatic juice enters the small intestine through the pancreatic duct and shifts the pH from acidic to basic final result: amino acids – done with protein digestion! Fat Digestion occurs in the small intestine fats don’t mix with water enzymes can digest only the fat molecules on the surface of the fat droplets fats are prepared for digestion by bile in the liver bile is a substance secreted by the liver and stored in the gallbladder bile emulsifies, or physically breaks down, large fat droplets into small fat droplets, increasing the surface area of the droplets available to the fat-digesting pancreatic enzymes bile doesn’t contain digestive enzymes lipase is the fat-digesting enzyme secreted in the pancreatic and intestinal juices and splits fats into 3 fatty acids and 1 glycerol final result: glycerol and fatty acids – done with lipid digestion! Absorption the end products of digestion are amino acids, simple sugars, fatty acids, and glycerol cells can absorb these small molecules through their cell membranes and use them for free energy and as raw materials for building cellular structures these small molecules pass through the cells lining the small intestine villi are the structures in the small intestine are responsible for absorbing the digested food villi: small fingerlike projections that increases the surface area of the intestinal lining each villus contains capillaries capillaries: tiny, thin-walled blood vessels that serve as entry points to the bloodstream simple sugars, amino acids, fatty acids, and some glycerol, minerals, and vitamins pass through the cells of the villi and enter the bloodstream through capillaries the blood carries the products of digestion to the cells inside the cells, the molecules are either broken down further to yield energy or used to synthesize substances the organism needs for growth and repair ©SarahStudyGuides 13 Nutrients glucose is the one nutrient in almost all foods food that comes from animals has cholesterol food that comes from plants doesn’t have cholesterol (bun, fries, cookies, sprite, etc….) lots of calories come from fat to add fiber to your meal: add fruit and vegetables soft drinks aren’t a good nutritional choice because they are mostly made of lots of calories and different sugars – they don’t have any other nutrients Seeds plant seeds can germinate in the dark because they break down amylose stored in the seed and can therefore carry out the process of cellular respiration how energy growth can come from a seed: enzymes amylose + H2O hydrolysis ATP enzymes glucoses + O2 H2O + CO2 + heat energy + free energy usable energy for cell work cellular respiration it’s important for germinating seeds to have water because water is needed for the seeds to use the process of hydrolysis which is needed to break down amylose. The seed can then carry out cellular respiration boiling the seeds would denature the enzymes of the seeds so they wouldn’t work the seed couldn’t grow planting the seeds in freezing soil or warm soil would also denature the enzymes so they wouldn’t work the seed couldn’t grow so: 1st and 2nd laws of therodynamics plants capture solar energy from the sun and transfer it into chemical energy through the process of photosynthesis photosynthesis produces chemical energy in the form of glucose and oxygen this chemical energy is converted into free energy available to do work in the form of the molecule ATP however, only 10% is converted to free energy – 90% of the energy that is converted from chemical to free energy is lost as heat energy which increases the entropy of the universe pancreatic amylase- enzyme that helps digests carbs breaks starch into polysaccharides, maltose, and glucoses trypsin- enzyme that helps digests proteins breaks polypeptide bonds, producing amino acids from those polypeptides lipase- enzyme that helps digests fats breaks fats into fatty acids and glycerol ©SarahStudyGuides 14 The Digestive System mouth -teeth -tongue -salivary glands esophagus stomach *liver, gallbladder, and pancreas small intestine large intestine anus * = the two accessory organs Function Mouth -mostly water (oral cavity) -pH is almost neutral Teeth and tongue -physical digestion What is secreted It acts on: It produces: salivary amylase Amylose smaller polysaccharides or maltose Proteins polypeptides -teeth: chewing -tongue: allows saliva to come in contact with food; moves food around Salivary glands -chemical digestion -salivary amylase: begins breakdown of starch Esophagus -carries food to stomach None! Stomach -really low pH -gastrin -protein digestion -HCl -regulation of gastrin, HCl, and pepsin secretion -pepsin -secretion bile salts large fat droplets small fat droplets -trypsin -polypeptides -amino acids Liver (secretions to gallbladder) Gallbladder -storage (secretions to small intestine) -transport of bile Pancreas -completes digestion of carbs, lipids, (secretions to ©SarahStudyGuides 15 small intestine) proteins -lipase -fats -amylase -carbs -fatty acids and glycerol -maltose Small intestine - 1/3 digestion of proteins, fats and carbs -proteinases -peptides -amino acids -lipases -fats -fatty acids and glycerol hi -2/3 absorption (villi) -secretin -regulation of pancreatic secretions - glucose -carbohydrases Large intestine -complex sugars -reabsorption of water -collection of undigested wastes Anus -waste exit ©SarahStudyGuides