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Chapter 29 – How Plants Colonized Land - Four main groups of land plants: o ________________________ (mosses) o ________________________ (ferns) o ________________________ (conifers – ex. pines) o ________________________ (flowering plants) - Bryophytes → “_____________________”; (no water tubes) - Pteridophytes → “___________________” but _______________________ - Gymnosperms → “__________________” with ______________ - Angiosperms → “___________________” with ______________________________________ - Closest relative to land plants = green algae called ________________________ - Features of Land Plants: o o - BUT….many algae have some of this….SO… - UNIQUE FEATURES connect land plants and charophyceans: o o - Differences between land plants and close algae relatives (so ONLY land plants have these): o __________________ meristems o ______________________ embryos o ___________________ of generations o Sporangia produces ______________________ o Gametangia produces gametes Archegonia = Antheridia = o DETAILS → Apical meristems → Multicellular, dependent, embryos → 1 Alternation of generations → Gametophyte with haploid cells → gametophyte produces … Sporophyte with diploid cells → sporophyte is produced by mitotic division of the _______________________; sporophyte produces haploid spores by ____________________ o ALSO…..water conservation → Prevents desiccation - Except for bryophytes, land plants all have true roots, stems, leaves because of vascular tissues o Xylem → o Phloem → - Evolutionary connections between land plants and charophyceans: o Homologous chloroplasts, peroxisomes, cell walls o ________________________ (structure in cell division) o _______________________ sperm (algae and many plants) o Molecular systematic → chloroplast DNA and rRNA - Bryophytes o Three phyla: __________________________, ____________________________, _______________________________ (mosses closest to vascular plants) o ____________________________ most conspicuous/ dominant phase - Vascular Plants o Food transport (__________________) and water transport (___________________) o 2 o ___________________________ = dominant generation Pteridophytes (seedless vascular) → FERNS Sporophyte = dominant o Homosporous → o Heterosporous → 2 kinds of spores - Megaspores = Microspores = Produces sori → Other information: o Seedless vascular plants formed “__________________” during carboniferous period 3 Chapter 30 – Evolution of Seed Plants - Reproductive Adaptations of Seed Plants: o Reduction of gametophyte NOT ________________________ like in pterophyta (ferns) o Evolution of seed Spores → Seeds → o Evolution of Pollen - Pollen → No water requirement (no flagellated sperm) Seed Plants o Vascular plants that produce seeds o Recall: sporophyte = dominant phase (ex. pine trees = sporophytes) o Two clades of seed plants: Gymnosperms → Angiosperms → Most diverse and geographically widespread _________________ known species o ANGIOSPERMS = phylum ___________________________ Monocots → Leaves with ______________________ veins ______________________ group (single ancestor) Dicots → Leaves with _________________________ veins _____________ seed leaves 4 Success of angiosperms due to: Flowers: Angiosperm structure specialized for reproduction Specialized shoot with 4 circles of modified leaves: sepals, petals, stamens, carpals o Sepals → _____________ and ____________ are NOT involved in reproduction o Petals → o Stamens → Produce _______________________ → male gametophytes o Carpals → Also called ________________ Produce __________________________ → female gametophytes Stigma → Style → Ovary → Ovules/ seeds are protected by the ovary o Seed forms from _________________ o Fruit forms from ________________ Fruit: Angiosperm structure that protects dormant seeds and aids in dispersal Mature ovary (fruit) o o Wall of ovary becomes a ______________________ → Simple fruit → 5 Aggregate fruit → Multiple fruit → Humans produce edible fruits via selective breeding (ex. apple, orange, cereal grain) Angiosperm Life Cycle Heterosporous → o Male → o Female → Steps of the cycle: o Anthers produce microspores o Microspores form pollen Male o Ovules produce megaspores o Megaspores from embryo sac Female o Pollen → sticky stigma of carpal Although they can self pollinate, most ____________________________ Cotyledons → Double Fertilization Characteristic of angiosperms Creates _______________________ central cell Function→ Specifics → o 1 pollen grain = __ sperm o 1 sperm fertilizes the egg → ___ zygote o 1 sperm fuses with the 2 nuclei to form 3n __________________ which becomes the … Note→ o Monocots store most food in __________________________ (ex. corn) 6 o Dicots transfer nutrients from the endosperm to the _____________________ (ex. beans) Coevolution Relationship between animals and angiosperms o o Plants and Human Welfare Agriculture (food!) is based on ________________________ Plant diversity is non-renewable o Humans are extinguishing plants → o Ethical concerns → o Practical concerns → ECOSYSTEMS ARE LIVING TREASURES!! 7 Chapter 35 – Plant Structure and Growth - Angiosperms → 250,000 species; most diverse and widespread - Genes and Environment affect Structure o Long-term – o Short-term – - Ex. Plant Organs: roots, stems, leaves o Differences between monocots and dicots MONOCOTS DICOTS Embryos Leaf Venation Stems Roots Flowers o Resources are needed from soil AND air; therefore, we need: ___________ system ___________ system ***These two depend on each other *** o Roots → Function → Monocots = Dicots = Both may have ______________________ (increase surface area; take up much water) OR __________________________ (above ground) roots 8 o Shoots → ________________________ (leaf) ________________________ (flower) Modified shoots (tubers, bulbs, etc) can be mistaken for roots o Stem → _____________________ bud (angle at leaf/ stem) have potential to bud _________________________ bud – concentrated growth; this inhibits axillary buds → However, if terminal bud is removed (pruning) the axillary bud will start to grow o Leaves → - Structure → Some have special functions (pg. 724) Plant Tissues → Dermal, vascular, ground o Dermal – o Vascular – Transports between roots and shoots Xylem → o Tracheids → o Vessel elements → o Fig. 35.8 pg. 725 9 Phloem → o Sucrose and other organic molecules thru tubes formed by chains of cells → _______________________________ elements End → sieve plates that have pores Non-conducting ____________________________________ may assist (via plasmodesmata) o Ground – - Neither dermal nor vascular Function → In dicots, divided as: Pith → Cortex → Plant Cells → parenchyma, collenchyma, and sclerenchyma o 3 basic types of cells Each cell has structural adaptations in the protoplast (contents) or cell wall Plasmodesmata → Middle Lamella → Primary Wall → Secondary Wall → o Parenchyma Primary walls → Most lack ____________________________ walls “Typical Cell” → therefore they are the __________________ specialized (all cells start out as parenchyma) 10 PHOTOSYNTHETIC! Perform most metabolic functions Division → o Collenchyma Primary cell walls → Grouped together to support ______________________________________________ Lack __________________________ walls o Sclerenchyma Function as ____________________________________ in plants Thick ________________________________ walls with ____________________ More rigid than collenchyma Mature cells cannot elongate, so they are present in cells that have ________________ _________________________ - Plant Growth and Development o Major differences between plants and animals: Totipotency → o Growth (definition) → o Development (definition) → o Plant Life Cycles Annual → Biennial → Perennial → Germ → Flower → Seed → Death o Meristems → Pattern of growth depends on the locations of the meristems 2 main types of meristems: Apical Meristems – 11 o Location → Fig. 35.12 pg. 729 o _______________________ growth (length) → gives rise to primary plant body Lateral Meristems – o o ____________________________ growth (thickens) → o Found in ________________________ plants o Growth Vocabulary → PRIMARY GROWTH!! (definitions) Root cap → Zone of cell division → Quiescent center → Primary meristems: Protoderm → Procambium → Ground → Zone of elongation → Zone of maturation → Miscellaneous vocabulary: stomata, guard cells, mesophyll (palisades/ spongy) → KNOW THESE!!!! o Growth Vocabulary → SECONDARY GROWTH!! 2 lateral meristems: Vascular Cambium → makes … 12 Cork Cambium → makes … o Periderm → o Bark → o Plants have developmental plasticity: Influenced by environmental factors 3 processes are important in the development of plants → Morphogenesis → o Depends on pattern formation → This depends on positional information = signals of some kind (chemicals) that indicate each cells location within an embryonic structure; ex. polarity Differentiation → o Depends on control of __________________________________ → regulating transcription and translation Growth → o Both contribute to plant form o Plane of division (ex. asymmetrical) is an important determinant of form o _________________________ accounts for 90% of expansion → o Fig. 35.26 pg. 739 13 Chapter 36 – Plant Transport - Recall: transport proteins, selective channels, proton pumps, chemiosmosis, cotransport, aquaporins KNOW THESE DEFINITIONS!!! - Water Potential o Depends on ________________________________ (hyper, hypo) AND ____________________________________ o Water moves from _________________ water potential to _____________ water potential o Adding ____________________ = lowers water potential o Adding ____________________ = increases water potential Fig. 36.3 pg. 751 o Pure water = water potential = ____ - Plant Cell Compartments o Symplast → o Apoplast → - Transport in Plants o Bulk Flow → movement of water/ solutes by____________________________ o _________________________________ increase surface area to absorb water and nutrients Mycorrhizae → o Endodermis → o Casparian Strip → o Movement of xylem sap Xylem sap consists of … Moves by 2 forces: 1. __________________________ → (“push”) Minerals accumulate in the stele = ________________________ water potential therefore water flows IN from the ________________________, which yields a positive pressure that force fluid UP the xylem 14 Guttation → o (See Pic pg. 757) o Occurs at night or when humidity is ______________ NOT the major mechanism 2. _________________________________________________ (“pull”) Water lost from transpiration; therefore, water is drawn from other cells by osmosis Cohesion → Adhesion → This is the MAJOR mechanism of xylem sap! Due to Hbonding of water! o So you could say that xylem transport is “_______________________”!! o Control of Transpiration Benefit of transpiration → HOW stomata open and close → change shape! Shape change → Lots of K+ = Little K+ = Fig. 36.13 pg. 760 Stomata usually open at __________________ (decrease in transpiration) CAM Plants → 15 o Movement of Phloem Translocation → Usually “food” is the disaccharide ________________________ Sugar “source” → Sugar “sink” → Always goes from source → sink Requires ________________________ transport Moves by bulk flow which is driven by pressure 16 Chapter 38 – Plant Reproduction - Alternation of Generations o The plant life cycle goes through 2 “variations” or generations o Gametophyte → o Sporophyte Produces _____________________________________ by ________________ - Usually the dominant generation of ___________________________ Flower Structure o Non-reproductive parts: ________________________ and _______________________ o Reproductive: Male → Female → o Complete Flower → o Incomplete Flower → - Development of Pollen (male gametophyte) o In ________________________________ (pollen sacs) are diploid cells (called ______________________________________) that do meiosis to form haploid __________________________________________, which give rise to _________________ (male gametophyte) o Two cells: ________________________________ cell = produces sperm; _____________ cell = produces pollen tube o Pollen = ____ sperm - Development of Embryo Sac (female gametophyte) o Ovary → ____________________ → megasporocyte → goes thru _________________ forming 4 megaspores – only ___ survives 17 o One end of the embryo sac has 1 egg and 2 _____________________ (which are used for … . - ) Pollination o Definition → - Prevention of self-fertilization o Ensures egg/sperm from different parents (genetic variety!) o Possibilities: - Self-incompatibility → Double fertilization o Synchronizes ____________________________________ with _________________________ o 1 sperm + egg = o 1 sperm + 2 polar nuclei = - Ovary Development o After double fertilization, ovule develops into a ___________________ and ovary develops into ___________________________ o Embryo development → divides into 2 parts: o Enclosed in seed coat o Mature seed = dehydrated (dormant!) - When rehydrated, forms embryonic root (_______________________) From seed to seedling o Germination of seeds depends on imbibition → 18 o _____________________________________ emerges first → water! o Rest of process → Fig. 38.14 pg. 794 - Asexual Reproduction o Asexual reproduction a.k.a. ______________________________ reproduction: - Results in clones Can grow from one _______________________________ cell Fragmentation → Apomixis → No genetic diversity Cuttings → Test-tube cloning; can cross genes Plant Biotechnology o Humans do selective breeding for our benefit o Transgenic → o Much debate → political, economic, ethical issues 19