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Imagine a population of 200 birds, some red and some yellow. The red allele, R, is dominant over the yellow allele, r. There are 72 RR birds, 96 Rr birds, and 32 rr birds in the population. ◦ What are the genotypic frequencies in the population? ◦ What are the allelic frequencies in the population? ◦ Using the frequencies you calculated in b, what would Hardy-Weinberg predict the genotypic frequencies should be? What about allelic frequencies? Is this population in Hardy Weinberg equilibrium? RR = 72/200 = 0.36; Rr=96/200 = 0.48 ; rr= 32/200 = 0.16 R=72 + 72 + 96 = 240/400 = 0.6; r = 32 + 32 + 96 = 160/400 = 0.4 P=R= Red q= r = yellow 32/200= 0.16 = yellow = rr= q2 √0.16=√q2 q = 0.4 then p = 0.6 RR = p2 = (0.6)2 = 0.36 = 36.00% Rr = 2pq = 2(0.6)(0.4) = 0.48= 48.00% rr = q2 = (0.4)2 = 0.16 = 16.00% Ms. Kim H. Biology Evolution on a small scale Change in allele frequencies from one generation to the next A process that leads to a change in a species Natural selection Genetic Drift Gene Flow Mutation Sexual Selection Microevolution explains how populations evolve Origin of different species SPECIATION Extinction of species Evolution of major features Protective Coloring ◦ Camouflage and Mimicry Physiological Adaptations ◦ Reproductive/Hormonal Changes ◦ Color changes Behavioral Adaptations ◦ Courtship dances/ songs ◦ “Fighting” tactics Also arise in response to environmental pressures ◦ Temperature, Antibiotic/pesticide resistance Animals who have greater fitness survive in environment and live to reproduce Random changes in DNA (mutations) can lead to greater or less fitness ◦ Produced by Sexual Reproduction ◦ Allow for DIVERSITY in a Population Adaptations allow an organism to survive better in their environment Directional ◦ Extreme form favored by natural selection Stabilizing ◦ Middle form most successful Disruptive ◦ Two extreme forms successful in separate environments 1. Convergent evolution Analogous Structures organisms that are NOT closely related independently evolve similar traits as a result of having to adapt to similar environments. 2 species acquiring same characteristics from 2 different ancestral species Ex: Dolphins & fishes Ex: Wings of bees & bats 2. Divergent evolution 2 species gradually become different Often occurs when closely related species diversify to new habitats Formation of 2 descendent species from an ancestral species Ex: Darwin’s finches Type: Adaptive radiation Homologous structures Appearance of numerous species over a short period of time Adapted species from a common ancestor when they are introduced to new environmental opportunities Typically occurs when a few organisms make their way to new, usually distant areas OR mass extinctions occur, which open up new niches Dubautia laxa KAUAI 5.1 million years 1.3 million MOLOKAI years MAUI OAHU 3.7 LANAI million years Argyroxiphium sandwicense HAWAII 0.4 million years Dubautia waialealae Dubautia scabra Dubautia linearis What type of evolution? A.Divergent B.Convergent C.Coevolution What type of evolution? A. Divergent B. Convergent C. Coevolution 3. Coevolution 2 organisms evolve (change) in response to each other Insects and the flowers (ex: orchids) they pollinate 4. Parallel Evolution 2 species evolve independently of each other, maintaining similar traits Usually occurs between unrelated species (but similar ancestors) that do NOT occupy the same or similar habitats ◦ Ex: Eutherians (placental) and Marsupial mammals What type of evolution? A. Divergent B. Convergent C. Parallel Evolution Origin of new species How would we identify if a species is “new”? Two Basic Patterns of Evolutionary Change Anagenesis (phyletic evolution) ◦ transforms one species into another Cladogenesis (branching evolution) ◦ the splitting of a gene pool, giving rise to one or more new species Anagenesis Cladogenesis Gradualism ◦ Species change slowly (gradually) over time Punctuated Equilibrium ◦ Species can make rapid “leaps” in evolution Time Gradualism model Punctuated equilibrium model Species is a Latin word meaning “kind” or “appearance” ◦ A population of organisms that produces viable fertile offspring in nature. ◦ They can NOT interbreed with other populations CASE 1 A mule is the offspring of a female horse and a male donkey. In contrast, the hinney is the offspring of a male horse and female donkey. The mule is easier to breed and larger in size than the hinney. For these reasons, the mule became an important domesticated animal. Horses have 64 chromosomes, donkeys have 62. The mule has 63 and cannot evenly divide, that is why the animal is sterile. CASE 2 A liger is a cross between a female tiger and a male lion. In contrast, the tigon is a cross between a male tiger and a female lion. These two species do not breed in nature because their habitats are so different. Lions live in open grasslands while tigers live in forests. In captivity, it is possible to produce ligers and tigons. Male ligors are sterile, but female ligers are fertile and may reproduce with either tigers or lions. CASE 3 E.coli is a bacterium normally found in the intestines. It is harmless and may actually be beneficial to the human digestive system. There is a pathogenic strain of E.coli that produces a toxin that can kill its human host. The two strains look very similar under a microscope. Comparison of their genomes reveals that the pathogenic strain lacks 528 genes found in the normal strain and has 1,387 genes not found in the normal strain. 5 Types: Morphological Group of individuals sharing similar characteristics Recognition Behavior/chemical recognition between individuals Genetic Range of variation in DNA- similar in individuals Cladistic Species defined as a branch in a cladogram Biological Group of individuals capable of interbreeding Similarity between different species different behaviors and songs Diversity within a species defined by capacity to interbreed. Speciation Speciation = the origin of new species ◦ Must explain how new species originate and how populations evolve Microevolution (genotype evolution) ◦ adaptations that evolve within a population’s gene pool Macroevolution (phenotype evolution) ◦ refers to evolutionary change at the population level ◦ Major biological changes evident in the fossil record