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Species & Speciation Morphological Species Concept • Species are based on comparison and differences existing in the physical characteristic between organism Morphological Problems • Two organisms that appear to be different species may be the same species • Likewise, organism with similar characteristics may be place in the same taxonomic groups when they shouldn’t be (fish and whales) Evolution & Phylogeny • After the publication of On the Origin of Species in 1859 by Charles Darwin classification took on new meaning. • Phylogeny is a form of classification with the goal of representing the evolutionary relationships and history of living organisms Biological Species Concept • Species are groups of actually or potentially interbreeding natural populations that are reproductively isolated from other such groups. (Mayr, 1942) • A species is a group of individuals fully fertile, but barred from interbreeding with other similar groups by its physiological properties. (Dobzhansky, 1935) Speciation • Process by which: a) subpopulations of one species diverge becoming adapted to different environments and reproductively isolated b) descendants of a species become adapted to an environment different than the one in which it was originally adapted such that they are qualitatively distinct from their ancestor Allopatric Speciation • Mode of speciation in which the subpopulations diverge because they are ‘separated by a geographic barrier’ • Common barriers include water (oceans to rivers), landforms (canyons and mountains), climate (glacial periods), and habitats (deserts, tropics, etc…). Geographic Separation • Once populations of a species are geographically separated the genes that determine their characteristics can no longer be shared (gene flow stops). • If the environment in the different geographic regions is not the same, then each population may diverge in their characters through natural selection. • If these populations are segregated long enough the populations may become new species. Reproductive Isolation • Pre-Zygotic Mechanisms 1) Temporal 2) Behavioral 3) Mechanical 4) Ecological 5) Gametic Mortality • Post-Zygotic Mechanisms 1) Hybrid Inviability 2) Hybrid Sterility 3) Low Fitness Hybrids Continental Drift & Speciation • Alfred Wegener • In the 1930’s suggested that continents were (and are currently) moving relative to each other and that at some time in the past they were joined together forming a super continent Pangea Evidence for Continent Drift • Continental Geography • Paleontology and Climate • Past Glacial Events Continental Geography • The eastern coast of South America and the western coast of Africa appear to fit nicely together like pieces of a puzzle Paleontology & Climate • The ranges of fossil plant and animal specimens of similar age appear to correlate well across current continental boundaries • Tropical fossils exist in Antarctic sediment suggesting that this continent has not always existed in its current location Past Glacial Events • Mapping of the extent of glacial rock remains on the different continent suggests particular connections between the continents in the past Plate Tectonics • Continental Drift was not taken seriously by the geological community until a mechanism that could explain the movement of the continents could be developed • Technical developments associated with WWII allowed a better understanding of the Ocean Floor and subsequent development of the mechanism of Plate Tectonics in the 1970’s Evidence for Plate Tectonics • Ruggedness and Age of the Ocean Floor • Oceanic Ridges and the process of Sea Floor Spreading • Oceanic Trenches and the process of Mountain Formation, Volcanoes, and Earthquakes Ruggedness of the Ocean Floor • Through development of sonar and more extensive use of submarines a picture of a rugged sea floor emerged (previously it was thought that the ocean floor was flat) • In fact, the Earth’s largest feature is the Mid-Atlantic Ridge running down the middle of the Atlantic Ocean Oceanic Ridges • At ocean ridges new crust arises from magma within the Earth forcing the existing plate out while a new border for the plate is being made Magnetic Striping • As new magma cools magnetic crystals within it line up with the magnetic poles of the Earth • The north end facing north and the south end facing south • As magma in the Earth’s core circulates “magnetic or polar reversals” occur • Our current north and south flip such that the magnetic crystal line up opposite of what they would be normally. Oceanic Trenches • Where two plates come together one is subducted under the other forming a trench • Tension builds up in the plate laying on top while heat may generate magma below (Mts & Volcanoes) • Mapping of the distribution of Earthquakes across the globe correlates well with places were plates are coming together • http://pubs.usgs.gov/publications/text/ Information on Continental Drift and Plate Tectonics