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Evolution Evolution = change in a _________, _____________, or _________________________ over time Populations vary by the __________________________________from one generation to the next Traits are represented by ____________________ Alleles may modify _____________ (form or structure), _____________, or _____________ Evolution = changes in allele frequency over time 2 areas of evolutionary study exist: 1. _____________ describes how populations change from one generation to the next describes how new species originate focus is on _____________ 2. _____________describes changes in groups of related species covers broad periods of time determines relationships between species ___________________________: early theory of evolution (1809) 3 important ideas: 1. _______________________: body parts of organisms can develop with increased usage, while unused parts weaken 2. _______________________: body features acquired during the lifetime of an organism (muscle bulk) could be passed on to offspring. THIS IS ________________________! 3. _____________________________: offspring produced with changes, future generations will be different Charles Darwin (1860) published “__________________________”; proposed the idea of _________________________= “Survival of the Fittest” Darwinism = Natural selection is the driving force NEO–Darwinism = today’s view, incorporates __________________________ NATURAL SELECTION the differences in survival and reproduction among individuals in a population is due to ________________________________________ some ________________ provide better traits for coping with the ___________________ More successful individuals produce ________________ Superior traits = ________________ increase _____________f 1 Favorable traits are _____________(selection acts _____the trait) Unfavorable traits are _____________ (selection acts __________the trait) Darwin’s arguments for Natural Selection: 1. _____________________________________________________________ (Darwin calculated that 2 elephants would produce a population of 19 million after 750 years if all offspring survive!) 2. Population sizes _____________________________ 3. ___________________________________________ 4. Individuals __________________________________________________ 5. There is ___________________among the __________________________ 6. Variation is ______________________________ 7. Only the ____________________________________________survive 8. Evolution occurs as _____________________________________________ Result = _________________________________________________________ Evidence for evolution does exist and can be observed. The ________________________by which evolution occurs have been the cause for debate through the ages! Other beliefs for the origin of life are not based upon science: Natural Selection may act upon a population in a variety of ways… KINDS OF SELECTION: 1. _________________________________ = eliminates individuals that have ___________________________________________ Individuals with the most common traits are _____________________ Frequency of common traits is ____________________ Other trait variations are ____________________________ 2. _______________________________= favors traits that are at _______________________________________________________ Traits of the opposite extreme are ______________________ Favored traits become more __________________ Examples: Insecticide resistance, Industrial Melanism (peppered moth) 2 3. ___________________________ (Diversifying Selection) = occurs when the environment ________________________________________________ Examples: weeds in Nature are tall, in lawns short 4. _______________________= the differential mating of males (sometimes females) in a population Females choose superior males for mates (_____________________) Male Competition: strongest males ___________________ Leads to _________________________= differences in the ___________________of males and females This can lead to disruptive selection Insert fig 8-1 SOURCES OF VARIATION In order for Evolution to occur _____________________________________ CONSIDERABLE __________________________exists in all populations Variation arises from the following mechanisms: 1. _________________________= provide the raw material, can invent alleles that never existed before 2. _________________________= new combinations of alleles, genetic recombination ________________________– exchange of DNA between non sister chromatids ______________________________of homologous pairs Random joining of ____________________ 3. ___________________= 2 copies of each chromosome in a cell; recessive alleles can be hidden 4. ___________________= mating of unrelated partners 5. _____________________________ = maintaining different phenotypes in a population Heterozygote Advantage: _____________________________ __________________: superior quality of offspring resulting from 2 different strains of inbred plants ____________________________(Minority Advantage): when the least common phenotypes have the biggest advantage Not all variation has _____________________________________! Much is _______________________________ i.e. fingerprints in humans The ____________________often determines whether a variation is _____________or whether it has selective value. 3 Causes of Changes in Allele Frequencies _______________________________was Darwin’s mechanism for evolution With the understanding of______________, the picture becomes more clear Other factors can change allele frequency other than natural selection 1. __________________________: increase or decrease in allele frequency due to the impact of the environment 2. __________________________: Introduce new alleles that may provide a selective advantage; most however are deleterious (harmful) 3. __________________________: alleles introduced by emigration (leave) vs. immigration (enter) 4. __________________________ : random increase or decrease of alleles; the effect is strongest in small populations __________________________: Occurs when the allele frequencies in a group of migrating individuals are, by chance, not the same as their population of origin (Amish & Polydactyly) __________________________: occurs when a population undergoes a dramatic decrease in size; the small group left becomes vulnerable to genetic drift (environmental catastrophes) 5. __________________________: mates chosen based on particular traits similar to their own, choose nearby individuals __________________________: individuals mate with relatives __________________________: females choose males based upon attractive appearance, behavior etc. GENETIC EQUILIBRIUM Occurs when ______________________________remain constant from generation to generation This is known as _________________________________________ AT GENETIC EQUILIBIUM THERE IS NO ___________________________! In order for this to occur the following conditions must be met: 1. No ______________________________________ 2. Mutations _________________________________________ 3. The population must be __________________________________ (No gene flow) 4. The Population is _____________________(No Genetic Drift) 5. Mating is ___________________________ IF ANY OF THESE CONDITIONS IS NOT MET, EVOLUTION ______________________________________! 4 Genetic Equilibrium is determined by evaluating the following values: 1. Allele frequencies for each allele (p, q) 2. Frequencies of homozygotes (p2, q2) 3. Frequency of heterozygotes (pq + qp = 2pq) Also the following two equations hold: 1. p + q = 1 (all alleles sum to 100%) 2. p2 + 2pq + q2 = 1 (all individuals sum to 100%) Example Problem: Suppose a plant population consists of 84% plants with red flowers and 16% with white flowers. The red allele (R) is dominant over the white recessive allele (r). q2 = 0.16 = White flowered plants (rr trait) p2 + 2pq = 0.84 = red flowered plants (RR and Rr traits) To determine the allele frequency of the white flower allele, calculate q by finding the square root of q2 q = .016 = 0.4 Since p + q = 1, p must equal 0.6 You can also figure out the percentages of individuals with the homozygous dominant and heterozygous condition 2pq = (2)(0.6)(.4) = 0.48 or 48% = heterozygotes p = (0.6)(0.6) = 0.36 or 36% = homozygous dominant 2 Speciation: the formation of a new species, can occur by way of 3 processes 1. _____________________________________ = a population is divided by some geographic barrier preventing interbreeding Gene frequencies in each population ______________________ If gene pools diverge enough, interbreeding will not take place if the barrier is removed As a result, _________________________________________ 2. ______________________________________ = formation of a new species without the presence of a geographic barrier Balanced Polymorphism: ex. Camouflaged insects Hybridization: two distinctly different forms of a species mate and produce offspring along a geographic boundary; the hybrid eventually diverges from both parent populations 3. __________________________ = relatively rapid evolution of many species from a single ancestor (Darwin’s Finches 14 varieties) 5 insert fig 8-2 Maintaining Reproductive Isolation: mechanisms at work if no physical barrier exists Prezygotic Isolating Mechanisms - ______________________________________________________________ 1. ______________________________: species don’t encounter each other 2. ________________________________: species mate at different seasons or at different times of the day 3. ____________________________________: when a species does not recognize a potential mate because it does not perform the correct mating ritual, display the proper visual signals etc. 4. __________________________________: male and female genitalia are structurally incompatible 5. ___________________________________: male gametes don’t survive in the environment of the female gamete Postzygotic Isolation Mechanisms - ______________________________________________________________ 6. ________________________: zygote fails o develop properly and is aborted 7. _____________________________________: Functional adults but they are reproductively sterile (mule) 8. ____________________________________: offspring have reduced viability or fertility Patterns of Evolution 1. _______________________________: 2 or more species originating from a common ancestor 2. _______________________________: 2 unrelated species that share similar traits; occur because each species has independently adapted to similar ecological conditions (not due to shared ancestry) 3. _______________________________: 2 related species that have made similar evolutionary changes after their divergence from a common ancestor 4. ___________________________: evolution of one species in response to another (plants & insects, predator & prey) 5. ___________________________: patterns of evolution occurring over extended periods of geologic time _____________________ gradual accumulation of small changes ___________________________________ long periods of “status quo” followed by short periods of rapid evolution Insert fig 8-3 Insert fig 8-4 6 Evidence for Evolution 1. _____________________________ = similar anatomy in different organisms • ____________________________ = similar structures shared due to inheritance from a common ancestor (limb structure) • _______________________________ = traces of a structure that no longer serves any useful purpose (hip bones in snakes) • _________________________________ = similar in purpose but not derived from a common ancestor (wings in birds vs. insects) 2. ___________________________________= embryos of related organisms develop in similar ways as embryos (vertebrates - gill pouches) “_______________________________________”. This means that during an organisms embryological development they will go through stages that resemble the stages they went through during their evolutionary development. 3. _________________________________ = chemical processes (metabolism) -genetic code, organic compounds, dependence on water - 98% of DNA is shared between humans and chimps 4. ______________________________: Transitional fossil forms = show intermediate stages of change (horses) 5. _____________________ provides variation - mutations 6. ________________________________________ can cause change (Industrial Revolution - Peppered Moth) 7. ______________________________ = bell shaped curve showing variation in a species , few extremes (rabbit color ranges from light to dark) • Directional Selection = shift in one direction (dark coats) • Disruptive Selection = acts on middle of curve (predation of medium coats) • Stabilizing Selection = selection against the extremes in stable conditions 8. _______________________________: unrelated species living in similar environments but in different geographic areas look alike 7