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Transcript
 YUKON CONSERVATION DATA CENTRE RARE PLANT INFORMATION SHEETS  INTRODUCTION  Yukon is home to more than 1200 species of plants, many (around 54) of which are found nowhere else in Canada. A better understanding of species distribution, abundance, threats and trends is necessary to sustainably manage the pressures on the territory’s natural resources. The Yukon Conservation Data Centre (CDC) aims to increase general awareness of rare plants and to help professionals and amateurs to identify rare species.  HOW ARE PLANTS ASSESSED?  The Yukon CDC assigns a conservation rank to all wild species known to exist in Yukon, in accordance with NatureServe methodology (visit NatureServe’s website for more information on Natural Heritage Methodology). Yukon CDC also compiles Tracking Lists for groups of wild species using these ranks, as well as current available information. Tracked species are those that are of global conservation concern by NatureServe, and/or are of specific conservation concern in Yukon as identified by the Yukon CDC, and/or are federally listed under the federal Species At Risk Act. Detailed location and status data are accumulated and maintained, and known populations are monitored as resources permit. Tracked species’ statuses are reviewed as information becomes available, and ranks may be up‐ or down‐graded based on new information. The Yukon CDC continually reviews species for tracking, and formally twice per year following COSEWIC assessment periods.  HOW TO USE THESE SHEETS  These sheets are organized alphabetically by scientific name. Individual sheets, groups of species, or the entire set may be printed for field or office use. Sheets include basic information on taxonomy, conservation status, habitat, range, and distinguishing features, and photos and drawings where available to aid field identification. A glossary of selected terms can be found at the end of these pages. Because sheets will be updated as information becomes available, you should check the Yukon CDC website for new versions. Please contact the Yukon CDC if you want us to compile habitat‐ or region‐
specific lists, or for other data requests.  WHAT TO DO IF YOU FIND A RARE PLANT  If you encounter one of the tracked species or something you suspect may be one of these species while working or travelling on the land, please contact the Yukon CDC with your information. Vascular Plant Field Form (available on the website) will help you record information such as:  location (mark a GPS waypoint if possible),  date and time,  observer name,  population size (count or estimate the number of individuals if possible) and,  area of extent (e.g. did you find them all over the mountainside on only at one site) Take photographs, if you can, of the habitat, substrate, and the plant (flowering or fruiting parts, leaves, and stems). Submitted photo(s) can use used to verify or add information. With your permission, the Yukon CDC could also add photos to that plant’s ID sheet. Voucher specimens (collected plant material) are critical in rare plant work. But rare plant populations can often be very small, making voucher collection questionable. The surveyor must weigh the pros and cons of taking specimens. Full vouchers (entire plants) should only be taken if the population is large enough to withstand the loss of an individual. Consider the “1 in 20 rule”; only take one plant if there are 20 others. While good quality photos are usually sufficient for identification, there really is no substitute for a specimen for verifying identification. Most plants can be identified without taking the whole plant.  CONSERVATION RANKS  The NatureServe Global and Yukon (i.e. “sub‐national”) rank for each species is listed in the upper left section of the sheet. NatureServe ranks are determined at the territorial level by the Yukon CDC and at the Global level by NatureServe using a standard method, and are regularly reviewed. In many cases, where the species is not found outside of Yukon, the Global Rank will also be calculated by the Yukon CDC. Ranks are calculated based on several factors: Rarity (range extent, population size); Trends (in population size or range); and Threats. NatureServe ranks are calculated and maintained by CDCs in each region and are then compiled by NatureServe to calculate National and Global ranks for species. This information is used by species experts, in part, to determine COSEWIC and General Status of Wild Species conservation status in Canada. From NatureServe Explorer: The conservation status of a species or ecosystem is designated by a number from 1 to 5, preceded by a letter reflecting the appropriate geographic scale of the assessment (G = Global), N = National, and S = Subnational). The numbers have the following meaning: X = presumed extinct or extirpated 1 = critically imperiled H = possibly extinct or extirpated 2 = imperiled NR = status has not yet been assessed 3 = vulnerable 4 = apparently secure U = unrankable with present information 5 = secure For example, G1 would indicate that a species is critically imperiled across its entire range (i.e., globally). In this sense the species as a whole is regarded as being at very high risk of extinction. A rank of S3 would indicate the species is vulnerable and at moderate risk within a particular state or province, even though it may be more secure elsewhere. In some cases, a “range rank” (e.g. G2G3) is designated if there is not enough information to specify a rank. “T” in a rank indicates sub‐ or infra‐species.  HOW TO CONTACT THE YUKON CONSERVATION DATA CENTRE  For more information on these or other species of conservation concern, to report a sighting or submit data, or to request data for plants or animals of conservation concern contact: Mailing Address: Phone: 867‐667‐5331
Environment Yukon Toll‐free: 1‐800‐661‐0408 local 5331
Government of Yukon Fax: 867‐393‐6405 Box 2703 (V‐5N) Whitehorse, Yukon, Canada Y1A 2C6 Physical Address: Email: [email protected] Environment Yukon Website: www.env.gov.ca/cdc 10 Burns Road Whitehorse, Yukon arte_laci
Yukon Rare Plant Information Sheet • May, 2012
Siberian Wormwood
Artemisia laciniata
Composite Family (Asteraceae)
Synonyms: A. laciniatiformis; A. macrobotrys, A. tanacetifolia
Habitat: Grassy flats, riverbanks, and open wooded slopes
with Calamagrostis purpurascens, Artemisia frigida and
Populus tremuloides
DISTRIBUTION
Yukon: Central Yukon; known only from the Yukon River
from Carmacks to Ballarat Creek where locally common
North America: Yukon and Alaska to Eurasia
World: Amphiberingian
B. Bennett
© NRC Research Press
Distinguishing features: Blades of leaves 2-3 times
pinnately divided; leaves glandular dotted; leaf segments
spreading; those of the lower leaves obtuse. Corollas
usually reddish. Vegetative plants may be confused with
Tanacetum bipinnatum.
B. Bennett
Global rank: Possibly secure (G4)
Yukon rank: Vulnerable (S3)
B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
botr_alas
Yukon Rare Plant Information Sheet • May, 2012
Alaska Moonwort
Botrychium alaskense
Adder’s Tongue Family (Ophioglossaceae)
Habitat: Gravel
DISTRIBUTION
Yukon: Central and southwestern Yukon
North America: Yukon and Alaska
World: Western North America
Global rank: Apparently Secure (G4)
Yukon rank: Imperilled to Vulnerable (S2S3)
Distinguishing features: This species can be separated
from similar pinnate-pinnatifid species as follow (from
Farrar, 2006*)
1a. Upper pinna bases obtuse (angle > 90°) pinna apices
rounded; sporophore stalk equal to trophophore length;
sporophore pinnately branched...............................B. pinnatum
1b. Upper pinna bases acute (angle < 90°); pinna apices
angular; sporophore stalk shorter than trophophore length;
sporophore divided into three main branches ..............go to 2
2a. Trophophore outline triangular (equilateral) with basal
pinnae nearly as large as the central rachis and pinnae;
pinna pairs 3-4; pinnae narrowly ovate to
oblong..................................................................B. lanceolatum
G. Brunner
2b. Trophophore outline narrowly triangular to broadly
ovate; basal pinnae not disproportionately enlarged;
pinna pairs 5-6; pinnae ovate to elliptic..................B. alaskense
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G. Brunner
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
botr_line
Yukon Rare Plant Information Sheet • May, 2012
Narrow-leaved Moonwort
Botrychium lineare
Adder’s Tongue Family (Ophioglossaceae)
Habitat: Dry gravel benches
DISTRIBUTION
Yukon: Southwestern Yukon; known only from Kluane Lake
and near Minto
North America: Scattered throughout unglaciated areas in
Alaska, Yukon, Quebec and the southern United States
World: North America
B. Bennett
Distinguishing features: Botrychium alaskense has the
narrowest pinna span of all moonworts, only broadening
where they divide. At maturity, its sporophore stalk is
twice the length of the trophophore.
B. Bennett
Global rank: Possibly Critically Imperilled (G2?)
Yukon rank: Critically Imperilled (S1)
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B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
coll_parv
Yukon Rare Plant Information Sheet • May, 2012
Scamman’s Springbeauty
Claytonia scammaniana
Purslane Family (Portulacaceae)
Synonyms: C. arctica; C. porsildii; Montia scammaniana
Habitat: Alpine tundra, talus and scree slopes on
serpentine rivers
DISTRIBUTION
Yukon: Western Yukon north of 61°N
North America: Yukon and Alaska to Russia
World: Endemic to Yukon and Alaska
B. Bennett
© NRC Research Press
B. Bennett - YG
Global rank: Vulnerable to Apparently Secure (G3G4)
Yukon rank: Vulnerable (S3)
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B. Bennett -YG
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
cori_ocho
Yukon Rare Plant Information Sheet • May, 2012
Pale Comandra
Comandra umbellata ssp. pallida
Sandalwood Family (Santalaceae)
Synonyms: C. pallida
Habitat: Prairie grassland and dry open coniferous
woodland
DISTRIBUTION
Global rank: Secure (G5T5)
Yukon rank: Imperilled to Vulnerable (S2S3)
A. Ceska
Illustration by J.R. Janish courtesy of the
University of Washington Press
Distinguishing features: A very distinctive species with
only one close relative (Geocaulon lividum) from which it
can be separated by having terminal cymes or panicles;
fruit dry and nutlike.
B. Bennett
Yukon: Southwestern and west-central Yukon; known only
from Kluane and Aishihik lakes and Five Finger Rapids
(Yukon River)
North America: Disjunct to Yukon, not known in British
Columbia. Found from Alberta to Manitoba, southwestern
United States
World: North America
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S. Cannings
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
Yukon Rare Plant Information Sheet • May, 2012
Dawson Wallflower
Erysimum angustatum
Mustard Family (Brassicaceae)
Habitat: Steep, dry, south-facing rubble slopes; rocky
outcrops on steep, open-wooded slopes; dry grasslands
Distribution:
Illustration courtesy of NRC Press
Distinguishing features: Erysimum
angustatum is separated from other
Erysimum species by its small stature (≤
Malpighiaceous hairs
30 cm), bright yellow colour, large petals
(≥ 12mm), and by being densely covered
in malpighiaceous hairs (hairs attached to a surface by
their centres).
B. Bennett
Global rank: Imperilled (G2)
Yukon rank: Imperilled (S2)
Species at Risk Act rank: Not At Risk
Illustration M.G. Harris
Yukon: Central and southwest Yukon
North America: Alaska and Yukon
World: Endemic to Yukon and Alaska
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B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
Yukon Rare Plant Information Sheet • May, 2012
Prairie-smoke
Geum triflorum
Rose Family (Rosaceae)
Habitat: Dry prairie and grassland; grassy flats in openings
in dry forest; dry gravel and sandy soils
Distribution:
Yukon: Southeast and south-central Yukon; known only
from the Takhini Valley, Beaver River and near Minto
North America: Western and central North America
World: North America
B. Bennett
Illustration by J.R. Janish courtesy of the
University of Washington Press
Distinguishing features: Geum triflorum is separated from
other Geum species by flower colour (straw to pale purple)
and habitat (grasslands).
B. Bennett
Global rank: Secure (G5)
Yukon rank: Critically Imperilled (S1)
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B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
mala_palu
Yukon Rare Plant Information Sheet • May, 2012
Bristle-like Quillwort
Isoetes echinospora
Quillwort Family (Isoetaceae)
Synonyms: I. muricata var. braunii
Habitat: Silty lake or pond margins; often submerged
DISTRIBUTION
Yukon: Southern Yukon south of 62°N
North America: Throughout Canada and Alaska, and
western and northeastern United States
World: North America
J. Line - YG
© NRC Research Press
J. Pojar
Global rank: Secure (G5)
Yukon rank: Critically Imperilled (S1)
J. Line - YG
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
suae_calc
Yukon Rare Plant Information Sheet • May, 2012
Yukon Goldenweed
Nestotus macleanii
Composite Family (Asteraceae)
Synonyms: Haplopappus macleanii; Stenotus borealis;
Stenotus macleanii
Habitat: Calcareous, dry stony and shaley slopes
DISTRIBUTION
Yukon: South-central Yukon
North America: Yukon
World: Endemic to central Yukon
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B. Bennett
© Province of British Columbia
Distinguishing features: Densely caespitose to matted
perennial from a multiheaded caudex; stems woody;
leaves linear, acute, rigid, keeled, 0.5-1.0 cm long, ciliate,
forming whorls, mostly all basal. Flowers golden yellow,
pappus tawny.
B. Bennett
Global rank: Vulnerable (G3)
Yukon rank: Vulnerable (S3)
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S. Cannings
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
poly_lonc
Yukon Rare Plant Information Sheet • May, 2012
Yukon Woodroot
Podistera yukonensis
Parsley Family (Apiaceae)
Habitat: Talus slopes and cliff crevices
DISTRIBUTION
Yukon: West-central Yukon
North America: Yukon and Alaska
World: Endemic to Yukon and Alaska
Global rank: Imperilled (G2)
Yukon rank: Imperilled (S2)
Species at Risk Act rank: Under review
B. Bennett
B. Bennett
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B. Bennett
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Illustration L. Mennell - YG
© NRC Research Press
Distinguishing features: Differing from Podistera macounii
by having a many-headed caudex. Leaflets entire or with
basal ones pinnate or deeply cleft.
For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
Yukon Rare Plant Information Sheet • May, 2012
Spiral Ditchgrass, Widgeon-grass
Ruppia cirrhosa
Ditch-Grass Family (Ruppiaceae)
Synonyms: Ruppia spiralis; Ruppia maritima var. occidentalis;
Ruppia maritima var. spiralis; Ruppia occidentalis; Ruppia
cirrhosa ssp. occidentalis
Habitat: Shallow, alkaline waters; marshes, ponds, sloughs
Distribution:
Yukon: Central Yukon; known only from Needlerock Creek
North America: Western and central Canada, sporadically
in United States
World: Circumpolar
Global rank: Secure (G5)
Yukon rank: Critically Imperilled (S1)
© Province of British Columbia
Distinguishing features: Ruppia cirrhosa is a submerged,
freely branched aquatic bearing alternate linear leaves. It
is separated from Ruppia spiralis by having a peduncle with
more than 5 coils, longer than 30 mm in fruit.
B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
Yukon Rare Plant Information Sheet • May, 2012
Mount Sheldon R agwort
Senecio sheldonensis
Composite Family (Asteraceae)
Habitat: Turfy places in subalpine meadows. Wet to moist
meadows and forest openings in montane to alpine zones
Distribution:
Yukon: South and central Yukon
North America: Yukon, Northwest Territories, British
Columbia
World: Endemic to mountains in NW Canada.
B. Bennett
Illustration L. Mennell - YG
Distinguishing features: A tall Senecio of alpine meadows, it is
separated from S. triangularis by its smaller stature (30-60cm
tall), broadly lanceolate leaves (not triangular-hastate) and
margins that are shallowly sinuate or wavy denticulate (not
denticulate).
B. Bennett
Global rank: Imperilled to Vulnerable (G2G3)
Yukon rank: Imperilled to Vulnerable (S2S3)
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B. Bennett
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For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
stel_dicr
Yukon Rare Plant Information Sheet • May, 2012
Williams’ Catchfly
Silene williamsii
Pink Family (Caryophyllaceae)
Synonyms: S. menziesii ssp. williamsii; S. menziesii var.
williamsii
Habitat: Heaths, disturbed ground, river gravel and bluffs,
roadsides
DISTRIBUTION
Yukon: West-central Yukon
North America: Yukon and Alaska
World: Endemic to Yukon and Alaska
Global rank: Apparently Secure (G5T4)
Yukon rank: Imperilled to Vulnerable (S2S3)
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B. Bennett
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Alaska Natural Heritage Program
Illustration L. Mennell - YG
F. Baldwin
Distinguishing features: Similar to S. menziesii but has
narrower leaves, longer calyx (12 mm vs 5-7 mm) and
reddish brown seeds.
For more information, contact the
Yukon Conservation Data Centre at 867-667-3684 or [email protected].
SELECTED GLOSSARY ABAXIAL  On the side of a structure turned away from the main axis. ACAULESCENT  Stemless. ACUMINATE  Tapering to a narrow tip or concave point, the sides generally concavely narrowing, "long‐
pointed”. ADAXIAL  On the side of a structure, turned toward the main axis. APEX  Tip, point or end. AWN  A slender bristlelike appendage, usually at the apex of a structure. AXIL  The angle between a structure and the axis to which it is attached (e.g., the angle between a leaf and the stem). AXILLARY  Used in reference to structures attached in the axil or juncture of stem and leaf stalks. BASAL  At, or forming, the base. BEAK  A long, slender projection on a broader structure such as a fruit or seed. BIDENTATE  With two teeth, "2‐cleft". BIFURCATE  Divided into two forks or branches. BIPINNATIFID  Twice branched. CAESPITOSE  Growing in dense, low‐lying clumps, "tufted". CALLUS  A hard or firm thickening of a structure. In grasses, the thick, hardened basal portion of the lemma. CALYX  A collective name referring to the sepals, the outer whorl of the perianth. CAUDEX  The woody, thickened and persistent basal portion of an herbaceous perennial that gives rise to leaves and flowering stems each year, i.e. “stem‐base”. CAULINE  Of, or pertaining to, the stem, i.e. "stem leaves". CIRCUMSCISSILE  A capsule that dehisces transversely in a circular line, like the opening of a lid. CONNATE  Union or attachment of like parts (e.g., petals), "fused". CORM  A thickened, underground stem with papery leaves; a storage organ. CRENATE  Toothed with rounded, broad teeth, "round‐toothed". CULM  The flowering stem of grasses, sedges and rushes. CYME  An inflorescence in which all floral axes terminate in a single flower with the terminal flower, at the apex of the inflorescence, blooming first. DEFLEXED  Bent downward or backward. DENTICULATE  Finely dentate or serrate (with sharp, spreading teeth). DIOECIOUS  Male and female reproductive structures borne on different plants. DISARTICULATE  The separation of structures at maturity. FARINOSE  Covered with a white mealy substance. FILIFORM  "Threadlike", slender and cylindrical. FLORET  A small flower, particularly one (1) in a floral cluster; pertains also to the flower of a grass spikelet which includes the lemma, palea and reproductive structures. GLABROUS  “Smooth”, without hairs or glands. GLAND  A spot or structure that produces a sticky or greasy substance. GLANDULAR  Having glands. GLUMES  The two small bracts located at the base of a grass spikelet; they do not subtend flowers. (subtend: to be positioned directly or closely beneath something.) GLUTINOUS  Covered with a sticky, glue‐like resin. HASTATE  Shaped like the head of an arrow with the basal lobes flaring outward, "arrowhead‐shaped”. HISPID  Bearing long, rigid and very sharp bristles or "bristle‐like" hairs. HISPIDULOUS  Slightly hispid. INCISED  Cut sharply, deeply and irregularly, as in leaf margins. INDUSIUM (PL. INDUSIA)  A scale‐like outgrowth that covers and protects the cluster of sporangia located on the underside of a fern frond. INFLORESCENCE  The flowering cluster, categorized by the arrangement of flowers on the floral axis. INTERCOSTAL  Between ribs or veins. INVOLUCRE  A rosette of bracts subtending an inflorescence or head. INVOLUTE  Margins longitudinally in‐rolled, on the upper side, toward the center. LANCEOLATE  "Lance‐shaped"; longer than broad, widening above the base and tapering to the apex. LEMMA  The outermost of the two bracts that subtend the grass floret (See palea). LIGULATE  With a ligule. LIGULE  Flattened part of the ray corolla in the Asteraceae family; in many grasses and some sedges and rushes, a membranous or hair‐like appendage on the inner side of the leaf at the junction of the leaf blade and the sheath. MALPHIGIACEOUS  Hairs attached to a surface by their centers. OBOVATE  The reverse of ovate; attached at the narrow end, "egg‐shaped". OBTUSE  Blunt, rounded. OVATE  Shaped like an egg, attached at the broader end, "egg‐shaped". Term applied to plane surfaces. PALEA  The innermost of the 2 bracts that subtend the floret in grasses (see lemma). PANICLE  A branching inflorescence on which younger flowers are borne at the apex. PAPPUS  The modified outer perianth series crowning the ovary and achene in the Asteraceae; usually plumose, bristle‐like, or of scales. PEDICEL  The "stalk" of a single flower in an inflorescence, or of a spikelet in grasses. PEDUNCLE  The single supporting stalk of a flower or inflorescence. PERIANTH  A collective term for the calyx and corolla. PERIGYNIUM  A "pouched" bract surrounding the pistil or achene in Carex species; a modified, tubular bract. PETIOLES  Leaf stalk. PHYLLARY  One of the bracts below the flowerhead in the species of the Asteraceae family. PINNATE  Compound leaf with the leaflets on both sides of an elongated axis. PINNATIFID  Pinnately lobed. PISTILLATE  Flowers possessing pistils, but no functional stamens (i.e., a female flower). PUBESCENT  "Hairy". PULVINATE  Cushion‐ or mat‐like. RACHIS  The main stalk of an inflorescence or compound leaf, the "axis of inflorescence". RETRORSE  Directed backward or downward. (Retroverse.) SCAPE  A leafless flower stalk growing directly from the ground. SCAPOSE  Bearing a scape. SCARIOUS  Having chaffy, membranous, brown or "wax‐papery" edges. SERRATE  "Sharp‐toothed"; like a saw edge. SESSILE  Without a stalk, "unstalked". SILIQUE  A long slender fruit of the Brassicaceae family which splits open along the edges leaving seed attached to a central partition. SINUATE  A pronouncedly waved leaf margin, "wavy". SORUS (PL. SORI)  A cluster of spore producing structures (sporangia) on the underside of a fern frond. SPATULATE  Broad and rounded at the tips; narrowing to the base, "spoon‐shaped". SPIKELET  A small spike; in grasses, the primary inflorescence composed of two glumes and one or more florets. SPOROPHORE  A leaf bearing reproductive structures; generally used in reference to the spore‐bearing leaves or ferns. STAMINATE  Describing a male flower that contains one or more stamens but no functional pistils. STELLATE  "Star‐shaped"; having many rays radiating out from a common center. STIPULE  An appendage that maybe present on each side of the base of a leaf or petiole at its insertion point on the stem. STOLONIFEROUS  Having a stolon: an elongate, creeping, above‐ground horizontal stem that roots at the nodes. STYLE  A generally elongated stalk connecting the ovary and the stigma through which the pollen tube grows. SUBGLOBOSE  Somewhat or slightly spherical. SUBTEND  To be positioned directly or closely beneath something. TOMENTUM  A covering composed of short, thickly matted, woolly hairs. TROPHOPHORE  A vegetative leaf or frond, one that does not have reproductive structures attached. TUBERCLE  A small, rounded swelling. TURION  A small, scaly bud‐like offset on an underground stem. VELUTINUS (VELUTINOUS)  Having a velvety texture. VERTICILLATE  Having whorls. VILLOUS  With long and soft, un‐matted, hairs, "long‐hairy". VISCOUS (VISCID)  Glutinous, greasy, or sticky. References E‐flora BC, 2011, E‐Flora BC Glossary of Botanical Terms. In: Klinkenberg, Brian. (Editor) 2010. E‐Flora BC: Electronic Atlas of the Plants of British Columbia [eflora.bc.ca]. Lab for Advanced Spatial Analysis, Department of Geography, University of British Columbia, Vancouver. [15 June 2011] Botany.com, 2010, Plant & Flower Dictionary. In: Botany.com: Plant Encyclopedia to Identify Plants, Flowers, Trees & More. [http://www.botany.com/index.16.htm]. Demand Media. [15 June 2011] Charters, Michael, 2011, California Plant Names: Latin and Greek Meanings and Derivations [http://www.calflora.net/botanicalnames/botanicalterms.html]. [15 June 2011]