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AP Environmental Science: Populations Name: ________ Where does this fit? I. The Living World Ecosystem Structure Biological populations and communities; ecological niches; interactions among species; keystone species; species diversity and edge effects; Ecosystem Diversity Biodiversity; natural selection; evolution; Natural Ecosystem Change Species movement; II. Population A. Population Biology Concepts Population ecology; carrying capacity; reproductive strategies; survivorship Chapter 4 Learning Objectives: 1. Dynamics of Natural Populations: Explain the three model ways populations grow, and describe the graph that would illustrate each. 2. Limits on Populations: Explain factors that limit populations, including those that increase as populations become denser (such as predation and resource limitation) and factors that are unrelated to the population density. 3. Community Interactions: Define the types of interactions that can occur between species in a community and the effect of those interactions on each species. 4. Evolution as a Force for Change: Describe the major ideas in the theory of evolution, such as inheritance and natural selection, and list examples of adaptations organisms have for survival. 5. Implications for Human Management: Describe at least three ways in which humans alter features in the environment to change population growth patterns. Key Terms: population ecology population density birth rate (b) death rate (d) growth rate (r) natural increase dispersal immigration emigration biotic potential intrinsic rate of increase exponential population growth environmental resistance carrying capacity (K) survivorship density-dependent factor density-independent factor zero population growth demographics highly developed countries developed countries infant mortality rates developing countries moderately developed countries less developed countries (LDCs) doubling time replacement-level fertility total fertility rate demographic transition preindustrial stage transitional stage industrial stage postindustrial stage age structure age structure diagram population growth momentum Baby Boom Immigration and Nationality Act Immigration Reform and Control Act (IRCA) Chapter 4.1: Dynamics of Natural Populations The tale of the Golden Frog, Atelopus zeteki. What are the pressures that led to this organism almost becoming extinct? – habitat loss and overcollection for the pet trade – an introduced chytrid fungus deadly to amphibians http://commons.wikimedia.org/wiki/File:Panamanian_golden_frog_01_2012_BWI_00395_zoom .jpg Dynamics of Natural populations. Ecosystems and the organisms within them are constantly changing. These changes occur because living organisms within the ecosystem face varying stresses from both living and non-living factors in the environment. Living things must respond to those stresses and their response changes not only the affected population but many others that are tied in some way to the affected population. • • • Again, what is a Population: a group of members of the same species living in an area Community: populations of different species living together in an area Populations grow with births and immigration They decline with deaths and emigration Change in population number = (Births + Immigration) – (Deaths + Emigration) • Population growth is a change (rate) in population size = r • Equilibrium: (births + immigration) = (deaths + emigration) Often, population growth is not zero • Population growth rate: amount the population has changed divided by the time it had to change A. Population Growth Curves • Population growth curves graph how populations grow and are used to determine: how fast a population could grow the population size now and in the future There are three basic scenarios that any given population may find itself in at any given point in time: What are these? exponential increase, logistic growth, and constant growth. 1. Constant growth is virtually never seen in nature. See Figure 4-1. Constant population growth rate: adding a constant number of individuals over each time period is the simplest type of growth to model. Is not generally found in nature but it’s a good comparison to other growth patterns Population number at the start + (A constant * Time) = Population number at the end (Ex: start with 2 individuals and a constant growth rate of 2 per week: at the end of 24 weeks you would have 50 individuals) 2. Exponential Increase—J-curve growth. When does this occur? Ppt 7 growth at a constant rate of increase Elaborate on Exponential growth. • • the doubling time remains constant (Ex: it takes 2 days to go from 8 to 16 individuals, as well as from 1,000 to 2,000 individuals) • r (or rmax) = the number of offspring individuals can produce in a given time if resources are unlimited • the number of times you multiply e by itself Starting population x constant (e) multiplied by itself a given number of times = Ending population • Under unlimited conditions, organisms with a high r have rapid population growth • Such growth is called an “explosion” but it cannot continue indefinitely • The population continues to grow and then dies off • Carrying capacity (K): the maximum population of a species a habitat can support without being ruined. In exponential growth carrying is exceeded. 3. Logistic Growth—S curve growth. When does this occur? This occurs when a population is at equilibrium and the number of births + immigration = the number of deaths + emigration. Logistic growth: some process slows growth so it levels off near carrying capacity results in an S-shaped curve • As the population approaches K, growth slows the population remains steady and growth = 0 the maximum rate of population growth occurs halfway to K Starting population + (reproductive capacity (r) * population * a number showing how far the population is from K) = Ending population What growth do the curves below represent? Use the graph below to explain real life growth. • J-shaped explosions are often followed by crashes • J-shaped growth results from unusual disturbances introduction of a foreign species, the elimination of a predator a suddenly changed habitat, arrival in a new habitat • Other populations show an S-curve In populations controlled by complex relationships between species Cycles of lower and higher numbers around K B. Biotic Potential Versus Environmental Resistance Create a note Biotic potential is “the number of offspring (live births, eggs laid, or seeds or spores set in plants) that a species may produce under ideal conditions.” A species’ biotic potential remains constant despite environmental pressures. Species are not often allowed to reach their biotic potential because of environmental resistance. – Measured by r (the rate at which organisms reproduce) – Varies tremendously from less than 1 birth/year (some mammals) to millions/year (plants, invertebrates) • Recruitment: survival through early growth stages to become part of the breeding population Young must survive and reproduce to have any effect on population size See Figure 4-3. What is Environmental Resistance? —is “the combination of all the abiotic and biotic factors that may limit a population’s increase”. Resistance may increase or decrease depending on population size and it controls a population’s size. – Prevents unlimited population growth Biotic: predators, parasites, competitors, lack of food Abiotic: unusual temperatures, moisture, light, salinity, pH, lack of nutrients, fire • Environmental resistance can also lower reproduction loss of habitat, pollution, changed animal migration A stable population results from interactions between factors that increase and decrease population] Reproductive Strategies— R-strategists (r-selected species) “produce massive numbers of young, but then leave survival to the whims of nature”. – results in low recruitment – rapid reproduction, rapid movement, short life span – adapted to a rapidly changing environment – “boom-and-bust” populations – “weedy” or “opportunistic” species, usually small – a housefly, dandelion, and cockroach are examples On the other hand, K-strategists (K-selected species) “have a much lower reproductive rate (that is, a lower biotic potential), but then care for and protect the young until they can compete for resources with adult members of the population”. See Discussion Topic #2 and #3. – – – – – – care for and protect young live in a stable environment already populated by the species larger, longer lived, well-adapted to normal environmental fluctuations their populations fluctuate around carrying capacity also called equilibrial species an elephant and California condor are examples Create a table to compare r and K strategists. What is the Life history of an organism? – – – progression of changes in an organism’s life age at first reproduction, length of life, etc. visualized in a survivorship curve Explain the three types of survivorship curve! – – – – – I. Type I survivorship: low mortality in early life most live their full life span (e.g., humans) Type III survivorship: many offspring that die young few live to the end of their life (oysters, dandelions) Type II survivorship: intermediate survivorship pattern (squirrels, coral) Limits on Populations What is the difference between carrying capacity and actual population size? • • • What factors limit population size? Population density: number of individuals per area Density-dependent factor: increases with increased population density • Disease, predation, food shortages, competition for food and space • Environmental resistance increases mortality • The more dense a population is the more resistant factors affect growth rate. Density-independent factor: its effects are independent of the density of the population • Spring freeze, fire • Is not involved in maintaining population equilibrium • factors include an unusual heat wave or hard freeze, severe weather and natural disasters. If a particular limiting factor moves outside an organism’s range of tolerance, then the organism dies irrespective of how many individuals there are in the population Mortality in a population may be related to either density dependent factors (related to population size per given area) or density independent factors (not related to population size per given area). In either case, when a population faces stress that leads to mortality, a critical number of individuals in that population are necessary for that population to survive in an ecosystem. • • Regulating a population’s size Only density-dependent factors can regulate a population (keep it in equilibrium) • Top-down regulation: control of a population (species) by predation • Bottom-up regulation: control of a population occurs as a result of scarcity of a resource (i.e. food) Factors controlling a population determine the effects of adding or removing a species to an area • Removal of a species can even affect species that don’t directly interact with it What is the Critical Number of a population? — • the minimum population base allowing the survival and recovery of a population a pack of wolves, flock of birds, school of fish • The group is necessary to provide critical interactions between members protection and support • If a population falls below this number surviving members become more vulnerable breeding fails extinction is almost inevitable If the number of individuals in a population drops below the critical number, density independent factors become very important. It is very difficult to pre-determine what the critical number for a particular population is because of the complexity of ecosystems. What is the connection between biodiversity loss and humans? • Human activities are responsible for the decline and extinction of species Humans change habitats, introduce alien species, pollute, hunt, etc. • Human activities are not density dependent they can even intensify as numbers decline Differentiate between the terms critical number and carrying capacity. What is density dependence? Critical number is the minimum size of a population below which it will not reproduce. The survival of the species is unlikely below the critical number. The carrying capacity is a characteristic of an ecosystem and “is the upper limit to the population of any particular species that an ecosystem can support....” Therefore, the critical number is the smallest number of individuals within a species that is required for the species survival while the carrying capacity is the largest number of individuals within a species that an ecosystem can support. Carrying capacity is a characteristic of an ecosystem, while critical number is a characteristic of the species.