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3/24/2017 PHYLUM CNIDARIA Greek root “cnido” means “stinging nettle” Thin-walled, soft-bodied animals with stinging tentacles General characteristics About 11, 000 species including jellies, corals, sea anemones - All members are aquatic (most are marine) - United by stinging tentacles - Tissue level of organization - - - - More complex than sponges but very simple Have true tissues but few primitive organs Radial symmetry (as adults) Many are colonial (such as Portuguese Man O’ War) Most are sessile, some are planktonic, few are nektonic Body Forms Many Cnidarians are polymorphic which means they have two or more separate body forms with alternation between forms The same species has two distinct forms: 1 3/24/2017 POLYP – cylindrical, vase-shaped - mouth faces up - sessile - asexual reproduction stage -Corals (left), hydras, & sea anemones exist in polyp form as adults MEDUSA - bell-shaped - mouth down - free-swimming/planktonic - sexual reproduction stage - jellies & Portugese man-of-war as adults 2 3/24/2017 Cells and Tissues Cnidarians have 2 tissue layers (most animals have 3): epidermis – protective outer layer contains stinging cells gastrodermis – inner layer forms gastrovascular cavity- cells in this layer aid in digestion & circulation * Between the two layers of tissue is a thick jelly layer called mesoglea. Mesoglea is very thick in jellies, it’s really why they are so jelly-like. Meloglea is not true tissue. 3 3/24/2017 Movement lack true muscle cells, but have special epidermal cells that allow tentacle movement medusa can open and close like an umbrella, allowing it to move by “jet propulsion” when water is forced out of the medusa Feeding and Digestion - All are carnivores - Mouth is surrounded by tentacles – pull in water & capture food - Tentacles contain stinging cells called cnidocytes - Each cnidocyte has a coiled stinger called a nematocyst that discharges to either trap or immobilize prey (able to differentiate between animate and inanimate object, doesn’t just fire at anything). The nematocyst injects venom. - Tentacles guide paralyzed food into the mouth - Nutrients are distributed via diffusion, waste is expelled through mouth loaded Discharged 4 3/24/2017 Nervous System - the mesoglea contains a nerve net: a diffuse network of nerve cells and their extensions - Mostly used for coordination contractions in body - Sensory cells connect the epidermis and gastrodermis statocysts Simple sensory “organs” in the medusa: 1. statocysts are used for balance 2. ocelli (eye spots) detect light REPRODUCTION Asexual Reproduction Polyps produce new polyps or medusae by budding Colonial hydrozoan 5 3/24/2017 Sexual Reproduction Mature medusa release gametes into the water After fertilization, the zygote becomes a ciliated larvae that swims around Eventually, the larva attaches to a hard surface and grows into a polyp Classes: 6