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Bell Pettigrew Museum of Natural History Interpretative Panels Text: Dr Iain Matthews Design: Steve Smart & Cavan Convery A University of St Andrews Development Fund Project School of Biology http://biology.st-andrews.ac.uk 3:2 Cnidaria P hy l u m Body Plan: • Radially symmetrical with tentacles • Diploblastic • No enclosed body cavity • Central ‘gut’ space with mouth, no anus Cnidaria S u b p hy l u m Super class Class The phylum Cnidaria is a large and diverse group of radially symmetrical animals that, despite their simple body form, can achieve high levels of complexity. They have a long fossil record, stretching back over 700 million years, however they have never moved onto land and all members of the phylum are aquatic. The majority are marine, although there are a small number of freshwater species. Their basic tissue level organisation is not radically different from that of sponges; two layers of cells lie on either side of the gelatinous mesogloea, forming a tube that encloses a central coelenteron, which has only a single opening surrounded by tentacles. The tentacles are covered with specialised stinging cell organelles, the nematocysts, and are used for both offence and defence. When triggered, threads are fired out from the nematocyst and inject a toxin into the prey, which are then drawn back in for digestion. Gut Endoderm Ectoderm secretion of an external casing, the calyx. Those with a calyx of calcium, the corals, have created vast reefs of great ecological and geological significance. However the calyx is merely a hard casing and the true skeletal system of the cnidarians is the mesogloea and/or the water contained within the coelenteron. The cnidarians show marked structural and life style diversity. Forms include solitary polyps (e.g. anemones and hydras), colonial sessile polyps (e.g. corals and hydroids ), colonial floating forms (e.g. siphonophores ) and solitary floating medusae (e.g. jellyfish ). Classification within Cnidaria Subphylum: Medusozoa Cnidarians exist in two distinct body forms; either as polyps, such as anemones and corals, or medusae such as jellyfish. On this basis, two subphyla are classified: the Medusozoa, in which a medusa occurs in the life history and the Anthozoa in which a medusa stage is lacking. Some cnidarians, through repeated asexual reproduction, form large colonies often with the See specimen. Subphlyum: Anthozoa Class: Hydrozoa Order: Limnomedusae Order: Laingiomedusae Order: Narcomedusae Order: Trachymedusae Order: Actinulida Order: Anthoathecata Order: Leptothecata Order: Siphonophora Class: Alcyonaria Order: Protalcyonaria Order: Stolonifera Order: Telstacea Order: Gastraxonacea Order: Gorgonacea Order: Coenothecalia Order: Alcyonacea Order: Pennatulacea Class: Scyphozoa Order: Stauromedusae Order: Coronatae Order: Semaeostomeae Order: Rhizostomeae Class: Zoanatharia Order: Actiniaria Order: Zoanthinaria Order: Scleractinia Order: Corallimorpharia Order: Ptychodactiaria Order: Antipatharia Order: Ceriantharia Class: Cubozoa Order: Cubomedusae Jellyfish, anemones corals and hydroids Some anemones have a special relationship with certain fish (mostly clownfishes), which live in amongst the tentacles and secrete a special mucus to avoid being stung. Some scyphozoan jellyfish (e.g. the lion’s mane jellyfish, Cyanea capillata ) can reach over 2 metres in diameter and have tentacles 60 to 70 metres long. The sea wasp is one of the world’s most dangerous jellyfish. It kills several people each year, most commonly from the tropical waters off Australia. Witnesses report victims covered in metres of tentacles and if the sting is sufficient death occurs in about 15 minutes. The cnidarian corals are responsible for building the huge reefs such as Australia’s Great Barrier Reef.