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Managing change in the uplands Mariecia Fraser I nt ro d u c t i o n Land with Less Favoured Area (LFA) status accounts for around 45% of the agricultural area of the UK, while for Wales this percentage rises to almost 80%. The traditional basis for farming these LFA areas is the raising of sheep and beef cattle, with the UK hills and uplands carrying around 12 million breeding ewes and more than a million suckler cows,in each case just over 60% of the UK total. Incomes on hill farms have always been low,with producers heavily reliant on government support and subsidies. In the decades following the Second World War, great efforts were made to improve agricultural productivity in the uplands. Large areas of hill land were set to the plough, and new swards of cultivated grasses specifically bred to cope with marginal conditions were established. Improved permanent pasture and temporary grass leys account for around one-third of the grazed areas of the hills and uplands today. Crucially, herbage production from these improved swards can be over five times higher than that from the indigenous grasslands and, consequently, such pastures are vital to the viability of livestock farming in the hills. The remaining two-thirds of UK grazed uplands which remain unimproved include some of the country’s most stunning and evocative landscapes. However, these rugged and diverse spaces, so valued for their aesthetic and amenity value, are also the product of man’s intervention, and while they contain habitats that are recognised as having international importance from a nature conservation perspective, there has been a significant decline in both the condition and extent of many of these ecosystems as a result of inappropriate management. Figure 2.1: Heather moorland, an upland community with international conservation significance. Prior to reform of the Common Agricultural Policy (CAP) in 2003, support for livestock farming in the LFAs was based on headage payments. This encouraged overstocking, which not only resulted in overproduction of livestock but also the overgrazing of fragile grassland environments. The oversupply of upland sheep and cattle had a negative impact on the development and availability of the high quality livestock needed to restore UK and export markets, while the overgrazing led to ecological imbalance and environmental deterioration. In particular, heavy grazing by sheep has been linked to a reduction in heather (Calluna vulgaris) and other dwarf shrub species, and their replacement by the invasive grasses Molinia caerulea and 9 Nardus stricta. The loss of key plant species from these ecosystems impacts upon their habitat value, which, in turn, has implications for internationally important insect and bird populations. Innovative management strategies that deliver both economically and environmentally sustainable upland systems, being multi-functional in terms of farming, rural development and the environment, are therefore urgently required. While the policy drivers for the hills and uplands may have changed substantially over the course of the last half century,the requirement for sound scientific evidence on which to base new policies is consequently as great now as it has ever been. The role of the grazing animal Critical to the development and refinement of management guidelines for such areas is an improved understanding of plant/animal interactions within grazed ecosystems, and recently our research has focused on quantifying the economic and environmental impacts of alternative grazing options. There is evidence from studies of both wild and domestic herbivores that variations in grazing habits exist between different livestock species. In general, larger animals with higher body weights have a greater requirement for nutrients in order to maintain body function and support growth and production. Crucially, this increase is not linear and, consequently, a larger grazing animal will have less time to spend selectively foraging for food, per unit of nutrition consumed, than a smaller animal. As a result, cattle are generally less selective in their feeding than sheep and will tend to focus their grazing on taller,less digestible patches of vegetation in order to maximise their intake. Furthermore, cattle are more likely to need to move to more accessible parts of the sward or to graze other communities once preferred areas have been grazed too closely to allow intakes to be maintained. Extending this hypothesis, experimental work has shown that periods of grazing by cattle can be beneficial in terms of controlling invasive hill species such as Molinia and Nardus. Studies investigating the foraging behaviour of free-ranging herbivores grazing complex swards have demonstrated that differences in selectivity can lead to niche separation between animal species, and hence give opportunities for complementary pasture use. Thus, the presence of one species can be beneficial to the productivity of other species in particular areas. Previous experiments with sheep and beef steers grazing permanent pasture have shown that both mixed and sequential grazing systems have the potential to deliver improved sheep performance without compromising cattle 10 productivity,leading to a higher total output per unit area. These benefits appear to arise from a combination of different preferences for plant species and parts, a reduction in sward rejection due to dung contamination, and a lower intestinal parasite burden. These findings demonstrate that, through changes to grazing management, it is possible to increase productivity without increasing inputs, thereby improving both economic and environmental sustainability. This work is now being extended to suckler-based beef systems, as the grazing behaviour of mature cows is known to be different to that of growing stock. On-going work is also clarifying the effect of adjusting the cattle to sheep ratio within mixed grazing systems, since national statistics reveal that there is considerable regional variation in the ratios of these two species. There is potential to extend this species comparison work to assess the impact of alternative livestock species and to explore the associated opportunities. In Wales, there is particular interest in understanding the potential role of hill ponies as grazing management ‘tools’. There is also an urgent requirement for improved knowledge of the ways in which interactions between animal species, diet selection and plant chemistry influence diffuse pollution from grazing systems, in order to deliver the biodiversity benefits of grazing while minimising nutrient losses to soil, water and air. Exploring breed differences In recent times, there has been a growing number of schemes encouraging the use of ‘rare’ or ‘at risk’ breeds as conservation grazers. While evidence in support of claims that grazing by these animals is beneficial in terms of improving biodiversity is largely anecdotal, it remains possible that differences in body size could give rise to breed differences in feeding behaviour. To explore this issue further,a series of grazing experiments with continental-cross and traditional beef steers investigated genotype x environment interactions when grazing improved permanent pasture and Molinia-dominated rough grazing. These studies established that utilisation of Molinia caerulea by the continental-cross and traditional breed steers was broadly similar. Performance results indicate that, while genotype may influence the liveweight gain of younger calves, breed type has little impact on the performance of older steers. In contrast, the type of sward grazed was found to have a highly significant effect on growth rates regardless of age (Figure 2.2). Likewise, while breed was found to have comparatively little effect on meat colour or stability, pasture type had highly significant effects on both. Pasture type also had greater impact on the fatty acid composition of the meat, but there was no effect on meat flavour. Figure 2.2: Effect of pasture type and genotype on the growth of finishing steers. More recently, detailed preference experiments with different breeds of sheep and cattle grazing heather moorland have been conducted. The results highlight the substantial difference in diet composition between cattle and sheep when grazing this type of vegetation (Figure.2.3) and suggest that sheep exhibit a greater degree of individual variation in grazing behaviour. Significant differences were identified in the proportions of key plant groups consumed by the two sheep breeds, with the Scottish Blackface diet containing a greater proportion of heather. These results demonstrate that important differences in forage selection exist between commercial breeds of hill sheep, and imply that there is potential for specific environmental goals to be met more effectively without compromising returns from conventional production systems. Given the wide regional variation in hill and upland sheep breeds across the UK,there is a pressing need for this work to be extended. Information on the comparative grazing behaviour of crossbred and composite sheep breeds is also required, given recent predicted changes to the structure of the sheep industry. Although the forage selected and consumed by the two breeds of sheep were different, the diets selected by the modern and traditional breeds of suckler cow were found to be broadly similar. These findings,along with those from the Molinia studies outlined above, imply that commercial breeds of cattle could potentially deliver many of the environmental benefits of grazing currently associated with traditional breeds. However, the continental cattle failed to thrive at exposed, high altitude sites. Thus, although the impact on the vegetation of such animals would be similar, there are likely to be production, and possibly welfare, implications associated with using this type of stock in very marginal areas. On-going research is determining the economic and environmental impacts of grazing options which integrate different sward types, and the longer-term performance of suckler systems based on Limousin-cross and Belted Galloway cattle. Figure 2.3: Principal component analysis of faecal concentrations of n-alkanes and long-chain fatty alcohols sampled in September from animals grazing heathland with 60% cover of Calluna vulgaris. 11 New challenges Looking to the future, climate change will impose further demands on upland systems. It has already brought greater awareness of the role the uplands play in providing ‘ecosystem services’ which help maintain a well-balanced, regulated environment. This has raised questions with regards to how to define ‘public goods’, how to pay for them, and how to balance these public goods with the need for food production. Uplands provide 70% of the UK’s water supply, and management in these areas strongly influences water quality. Predicted changes to rainfall pattern will lead to upland areas playing an even greaterrole in water management and flood control for low-lying areas downstream. At the same time, temperature rises are predicted to be greater at higher altitudes, and this, together with a reduction in summer rainfall, will increase the risk of summer drought and wildfires in hill and upland areas. However, such temperature changes and an associated lengthening of the growing season could enable the uplands to become agriculturally more productive. Such changes may lead to the adoption of new crops and livestock breeds in the uplands, potentially compensating for a loss of food production in lowland areas associated with the growing of bioenergy crops. The uplands could therefore play a crucial role in ensuring food security, but this capacity is as yet largely undefined and untested. There is also a pressing need for more information regarding the effect of agricultural land abandonment in the hills and uplands, notably on landscape, ecosystems, water retention, carbon balances and biodiversity. With so many policies and support schemes relating to the uplands currently under review, perhaps the greatest challenge of all will be agreeing an evidence-based vision for these areas and developing an integrated framework for delivery. Mariecia Fraser email: [email protected] Figure 2.4: Belted Galloway cattle grazing Molinia-dominated rough grazing at Bronydd Mawr. 12