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Origin and History of Life Where it came from and how it got here Chapter Outline Primitive Earth Origin of First Cells Fossils The Precambrian The Paleozoic The Mesozoic The Cenozoic Continental Drift Mass Extinctions Beginnings The Universe (maybe) Primitive Earth—Not paradise as we know it—much different than Earth today Simple to complex Question: How could these simple molecules be made into more complex ones? Answer: Abundant energy sources, lots of time. Origin of organic molecules Alexander Oparin (1938) — a Russian biochemist with vision Stanley Miller (1953) – tested the hypothesis Miller Apparatus Other models Carbonaceous Panspermia chondrites Panspermia Life brought in on asteroids and comets EX. Deinococcus radiodurans Abiotic Chemical Evolution Synthesis and accumulation Polymerization Aggregation Origin of heredity Origin of First Cells Keep in mind!!! Chemical natural selection Conditions of primitive earth Timescale Evolution of Macromolecules RNA first hypothesis Protein Clay first hypothesis hypothesis RNA first hypothesis RNA contains information like DNA RNA is catalytic-- -- ribozymes. Protein First Hypothesis Most enzymes are proteins Proteinoids Proteinoid microspheres Faithful replication? Clay hypothesis Unique chemical properties of clay. Combines parts of the RNA first and protein first ideas. Protocell Evolves Protocells -- precursors to the first true cells. lipid-protein membrane Proteinoid microspheres Coacervates Liposomes Protocell Anatomy Energy Management Were the first protocells heterotrophs or autotrophs? Origin of First Cells The Origin of Life 2 Fossils Fossils are remains and traces of past life Why are they so rare? Fossils Dating Fossils Relative Dating Absolute dating Combination Strata The Precambrian Age of microbes 570 mya - 4.6 bya The First Cells —3.5-4 billion years ago What were they like? Energy management? Aquatic or terrestrial? Prokaryote or eukaryote? Evolution of Autotrophs Life as Energy Management Chemoautotrophs Photoautotrophs Atmospheric O2 ~2 BYA Atmospheric O2—SO WHAT? O2 is very reactive—Poison. Cells had to deal with this Aerobic Anaerobic Facultative anaerobes Changes in atmosphere– O2 and ozone. So? Consequences for energy acquisition Circle of Life Energy Respiration Carbon compounds, O2 CO2, H2O Photosynthesis Energy Eukaryotes evolve More complex Origin? Endosymbiont hypothesis nucleoid (DNA) ribosomes food granule prokaryotic flagellum plasma membrane cytoplasm cell wall nuclear pore chromatin (DNA) nucleus nucleolus nuclear envelope flagellum rough endoplasmic reticulum cytoplasm intermediate filaments plasma membrane ribosome lysosome microtubules smooth endoplasmic reticulum Golgi complex free ribosome vesicle mitochondrion vesicle 1. Anaerobic, predatory prokaryotic cell engulfs an aerobic bacterium. aerobic bacterium 2. Descendants of engulfed bacterium evolve into mitochondria. 3. Mitochondria-containing cell engulfs a photosynthetic bacterium. 4. Descendants of photosynthetic bacterium evolve into chloroplasts. Multicellularity Evolves Examples? Volvox—Simple, multicellular Advantages? Summary of Precambrian Earth forms Prokaryotes evolved Photosynthesis Eukaryotes Multicellularity Sexual reproduction Paleozoic Paleozoic 2 Mass extinction---Disappearance of a large number of species within a relatively small time interval. opportunities for speciation PALEOZOIC- MAJOR EVENTS What were the major events of the Paleozoic? 1. three mass extinctions 2. Invertebrates dominated – exoskeleton 3. Primitive plants invaded land PALEOZOIC- MAJOR EVENTS 1. 2. Vertebrates appeared (jawless fishes) -- internal skeleton Insects and amphibians invaded the land -- -- evolution of flight, evolution of lungs, evolution of stronger limbs, strengthened internal skeleton PALEOZOIC- MAJOR EVENTS 1. 2. 3. Reptiles appear a) Shelled egg b) Scaly skin c) Reptiles diversify amphibians declined Coal forming forests Conifers appear Invasion of Land Plants Invertebrates Vertebrates Mesozoic MESOZOIC- MAJOR EVENTS The age of reptiles Angiosperms evolve-insects diversify Mammals and birds Evolution of endothermy QUESTIONS From what to did dinosaurs evolve? From what to do to mammals evolve? From what did birds evolve? What caused the mass extinction of dinosaurs? Were the dinosaurs endothermic? Endothermy vs Ectothermy Endothermy Warm –blooded Requires a high use of energy High metabolic rate Disadvantages? Ectothermy Cold blooded Less energy necessary Regulation? behavioral structural inertial ectothermy Cretaceous Mass Extinction Bolide theory -- -- the most widely accepted theory of the Cretaceous Mass extinction. Bolide -- -- and exploding asteroid. Cenozoic CENOZOIC- MAJOR EVENTS Three mass extinctions Decline of reptiles Diversification of Angiosperms Insects Birds Mammals Factors That Influence Evolution Continental Drift Positions of continents and oceans are not fixed. How does this explain the presence of marsupials and monotremes in Australia? Continental Drift Continental Drift Plate Tectonics Movements of Earth’s crust which is fragmented into slablike plates that float on a lower hot mantle layer. Plate Tectonics Mass extinctions A mass extinction occurs when a large fraction of all living species becomes rapidly extinct. The fossil record shows that at least five major mass extinctions have occurred in the past 500 million years. Impacts of asteroids on Earth are suspected as a primary cause of mass extinction. Mass Extinctions Mass extinctions have occurred throughout history. At the ends of: Ordovician Continental Drift Devonian Bolide Event Loss of 70% of marine invertebrates Mass Extinctions Permian Excess carbon dioxide. Loss of 90% of ocean species. Triassic Meteorite Collision Cretaceous—probably a bolide. Permian Mass Extinction At the end of the Permian period (about 245 mya), destroyed 90-95% of marine and 70% of land species. Possibilities? The glaciation of Gondwanaland the increased fluctuation of global temperatures volcanic eruptions occurring in Siberia Mass Extinctions Intrinsic gradualism Extinction caused primarily by climate change or other gradual processes. EX. Global warming Extrinsic catastrophism Extinction caused by a catastrophe Ex. Asteroid collision with Earth Review Primitive Earth Origin of First Cells Fossils The Precambrian The Paleozoic The Mesozoic The Cenozoic Continental Drift Mass Extinctions