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ANIMAL, PLANT & SOIL SCIENCE E4-2 PLANT INTERNAL STRUCTURE AND GROWTH Interest Approach Take the students outside to see a small tree, bring a small tree to class, direct attention toward a small tree outside the window, or show an image of a small tree. Identify a small lower limb on the tree. Then, ask the students where that limb will be in 50 years if the tree continues to grow. Students often assume that a tree grows from the ground and that it begins by enlarging at the ground level; they fail to realize that growth occurs from meristem tissues. Objectives 1 Review the functions of plant cell components. 2 Examine the role of cell division, cell elongation, and cell differentiation in plant growth and the different types of plant tissue. 3 Identify and describe the basic types of plant cells. 4 Compare and contrast primary and secondary plant growth. Terms _ apical meristem _ bud primordia _ cell _ cell differentiation _ cell division _ cell elongation _ cell wall _ cellulose _ chlorophyll _ chloroplasts _ chromatin _ chromosomes _ collenchyma cells _ companion cells _ cork cambium _ cork cells _ cytoplasm _ dermal tissues Terms _ endoplasmic reticulum _ fiber _ genes _ golgi complex _ ground tissue _ lateral meristem _ leaf primordia _ lignin _ meristems _ mitochondria _ nucleolus _ nucleus _ organelles _ parenchyma cells _ periderm _ phloem _ pith _ plasma membrane Terms _ plastids _ primary plant growth _ ribosomes _ rough endoplasmic reticulum _ sclereid _ sclerenchyma cells _ secondary plant growth _ sieve tube member _ smooth endoplasmic reticulum _ tracheid cells _ vacuole _ vascular cambium _ vascular tissues _ vessel elements _ xylem What are the functions of plant cell components? I. The cell is the most basic unit of life. It is the smallest unit that can perform functions of life. Plant cells are composed of the following items. A. The cell wall is made of multiple layers of cellulose, which is a polysaccharide or complex sugar molecule. The layers of cellulose offer great strength. Therefore, it is no surprise that cellulose is the primary ingredient of wood. Once a cell has stopped growing, the cell wall thickens and becomes rigid. What are the functions of plant cell components? B. The feature that really defines a cell is the plasma membrane or cell membrane, which is a thin sheet that holds the contents of the cell together and serves as a protective barrier from the surrounding environment. The fluid mosaic model of the plasma membrane consists of a lipid bi-layer in which proteins are imbedded. While the plasma membrane restricts the movement of some materials, it allows and even initiates the movement of other materials into and out of the cell. What are the functions of plant cell components? C. Cytoplasm is the living semi-fluid material inside the cell membrane and is home to a number of specialized structures called organelles. D. Organelles are small structures inside the cell that carry out the physiological processes of the organism. What are the functions of plant cell components? 1. Plastids are organelles that produce and store food. a. Chloroplasts are a type of plastid that contains green pigments called chlorophyll. Chlorophyll traps light energy for photosynthesis. Chloroplasts are found in cells exposed to light, so they are abundant in leaves and absent from root cells. b. Chromoplasts are plastids that give flowers and fruits their color. What are the functions of plant cell components? 2. The endoplasmic reticulum (ER) is an internal network of membranes extending throughout the cytoplasm. The endoplasmic reticulum contains many types of enzymes that catalyze different types of chemical reactions. There are two distinct forms of endoplasmic reticulum: smooth and rough. a. The smooth endoplasmic reticulum is the site for the production of lipids (fats) and hormones. It is also a site that breaks down toxic chemicals. b. The rough endoplasmic reticulum produces the proteins for the cell secretion and cell membranes. What are the functions of plant cell components? 3. Proteins are processed, sorted, or modified in the golgi complex. These processes result in the complex molecules needed for plant growth. Many proteins manufactured in the ER pass through the golgi complex. 4. Mitochondria convert sugar into energy through cellular respiration. With mitochondria, the plant is able to convert the food that it has made and stored into a form of energy. What are the functions of plant cell components? 5. The nucleus is the command center of the cell. It holds the genetic information for a cell’s activities within a nuclear membrane. a. The nucleus is surrounded by a nuclear membrane. Flowing through the nuclear membrane are materials needed for protein synthesis and fluids carrying chemical messages between the cytoplasm and the nucleus. b. Chromosomes contained in the nucleus hold the genetic blueprint of the organism and direct all functions of the cell, including protein synthesis. Chromosomes are made of chromatin. (1) Chromatin is a complex of deoxyribonucleic acid or DNA molecules (35 percent), RNA (5 percent), and protein (60 percent). (2) Segments of chromosomes, or genes, are units of hereditary data that dictate the activity and structure of a cell. Each chromosome contains thousands of genes. c. A nucleolus is a specialized structure in a nucleus that is the site of ribosome synthesis. What are the functions of plant cell components? 6. Ribosomes are bead-like structures found on the endoplasmic reticulum. Ribosomes are the major site for the assembly of amino acids into proteins. 7. The vacuole is a large sac bound by a membrane. It may occupy up to 90 percent of the cell. It serves to transport and store water, foods, salts, minerals, pigments, proteins, and wastes. What are the functions of plant cell components? E. The organelles within a cell have thin membranes used to compartmentalize or isolate different conditions from other organelles. For example, mitochondria and chloroplasts are bound with membranes. Different intracellular pH, enzyme systems, and more can be noted within each organelle. This enables the cell to perform different metabolic activities at the same time. What is the role of cell division, cell elongation, and cell differentiation in plant growth? What are the different types of plant tissue? II. Growth of higher plants takes place through three essential processes, and vascular plants have three types of mature tissues. A. Cell division is simply the increase in the number of cells, which is accomplished through mitosis. B. Cell elongation is the enlargement of the individual cells, which results in growth of the entire tissue or organ. Auxins and gibberellins are responsible for cell elongation. What is the role of cell division, cell elongation, and cell differentiation in plant growth? What are the different types of plant tissue? C. Cell differentiation allows cells to take on specific functions. Cell differentiation is important to multicellular plants, whereas single-cell plants have no need to have different cell types. The cells of the higher plants are programmed to become specialized and make up the various plant organs. Some cells specialize as root cells that absorb water and nutrients. Some conduct water and minerals throughout the plant. Others become leaf cells containing chloroplasts and function to produce food for the plant. What is the role of cell division, cell elongation, and cell differentiation in plant growth? What are the different types of plant tissue? D. Vascular plants have three types of mature tissues. 1. Dermal tissues consist of the epidermis or external covering of the leaves, stems, and roots. Dermal tissues offer a barrier to infectious organisms and other invaders. a. The epidermis is made of closely packed cells that secrete a waxy cuticle to reduce water loss. The epidermis is a single layer of cells associated with plants that exhibit primary growth. Specialized epidermal cells called guard cells control the opening and closing of the stomata through which the diffusion of carbon dioxide, oxygen, and water vapor is allowed. b. Dermal tissue on secondary woody growth is a few too many cells thick and forms the familiar bark of trees. What is the role of cell division, cell elongation, and cell differentiation in plant growth? What are the different types of plant tissue? 2. Vascular tissues are the conductive vessels of the plant. They transport water, minerals, food, and hormones. Specialized vascular tissues include the xylem, which conducts water and minerals upward from the roots, and the phloem, which transports food. 3. Ground tissue (fundamental tissue) includes all the parts of the plant other than dermal or vascular tissues. The bulk of a plant consists of ground tissue consisting of parenchyma, collenchyma, and sclerenchyma cells. What are the basic types of plant cells? III. Genetic information and environmental conditions determine the types of cells that a plant produces. With the appropriate signals, various types of cells are produced. A. Parenchyma cells are found throughout a plant and typically lack a secondary cell wall associated with rigid strength for support. The palisade layer and spongy mesophyll cells in leaves are parenchyma cells. Modified parenchyma cells compose a large portion of the pith (the center part of a stem). There they serve as storage facilities for starch, oil, water, and salt. The pith is very evident in corn and sugar cane. What are the basic types of plant cells? B. Collenchyma cells are located under the epidermis of the stem, along leaf veins, and at corners of angular stems. They have thick, yet flexible cell walls that provide structural support for the plant. C. Sclerenchyma cells are found throughout a plant and function as a support. These cells have a hard, thick secondary cell wall composed, in part, of a material called lignin, which is a molecule that provides strength and rigidity to the cell. A long type of sclerenchyma cell is called fiber. Fiber cells are abundant in wood and bark. A form of sclerenchyma cell that is short and cubical is the sclereid. It is found in the shell of nuts and the stones of fruit, such as peaches. What are the basic types of plant cells? D. Tracheid cells are one of four types of cells found in the xylem. The others are vessel elements, parenchyma, and fibers. Tracheids are long and tapering in shape. They are dead cells and hollow. Their role is to conduct water and minerals throughout the plant. Water passes from one cell to another through thin places in the cell wall called pits. Tracheids also contribute to the structural support of the plant. What are the basic types of plant cells? E. The vessel elements are found in the xylem where they conduct water and minerals and provide structural support for the plant. Vessel element cells are hollow. At the end of the cell walls, there are perforations or holes that allow the free flow of water from one cell to the next. Vessel cells also have pits along the sides of the cells that permit the lateral movement of water. What are the basic types of plant cells? F. The sieve tube member is a major element of the phloem. Other cells that compose phloem tissue are companion cells, parenchyma, and fibers. Sieve tube members are stacked end to end to form sieve tubes. There are holes at the end of each cell called sieve plates through which sugars are moved. Sieve tube members are living cells at maturity, but many organelles—including the nucleus, vacuole, and ribosomes—vanish as the cell matures. What are the basic types of plant cells? G. Companion cells are associated with sieve tube members. They have many connections with adjacent sieve tube members. Companion cells do not conduct sugar throughout the plant. They do, however, play an important role in transporting sugars from photosynthetic parenchyma cells to the sieve tube members. What are the basic types of plant cells? H. Cork cells are produced to the outside of the cork cambium and result in the outer covering of secondary growth. Cork cells have waterproof characteristics and fire-retardant attributes. How do primary and secondary plant growth compare? IV. The growth of a plant takes place in specialized areas of the plant called the meristems. The meristem tissue consists of unspecialized cells that are capable of mitosis. Two types of growth result from cell division in meristem regions: primary and secondary growth. A. Primary plant growth occurs in areas called apical meristems. Apical meristem tissue is found at the tips of roots and at the end of stems. Growth at the apical meristem increases the length of the plant. Primary growth occurs in herbaceous and woody plants. How do primary and secondary plant growth compare? 1. The meristem of a root is located right behind the root cap, which is a layer of cells that protect the meristem as the root grows through the soil. Just behind the root cap is the area of cell division. Further back is an area of cell elongation. As the cells in the root mature, they differentiate to perform specific functions. In a root system, the cells might become root hairs, xylem, phloem, etc. 2. The meristem of the shoot has areas of cell division, cell elongation, and cell maturation. The structure of the stem apical meristem differs from the root apical meristem in that it has embryonic leaves known as leaf primordia and embryonic buds called bud primordia. As the cells in these immature structures divide, elongate, and mature, they become the leaves and stems of the plant. How do primary and secondary plant growth compare? B. Secondary plant growth takes place only in woody plants. It takes place in the lateral meristem; this tissue is found on the sides of roots and stems. Secondary plant growth involves cell division in layers ringing the stem. The result is an increase in the width of the stem or trunk of the plant. Two lateral meristems are involved in secondary growth. How do primary and secondary plant growth compare? 1. Vascular cambium is a layer of meristematic tissue found between the wood and the bark. It produces secondary xylem to the inside of the vascular cambium and secondary phloem to the outside. Cell division occurs only when the plant is actively growing, primarily during the spring and summer. 2. The cork cambium is located in the outer bark region, and the cells produced there form the periderm (the outer bark). Cork cambium produces cork cells that replace the epidermis present during primary growth. The periderm ranges from several cells in thickness to many cells. REVIEW 1. What are the functions of plant cell components? 2. What is the role of cell division, cell elongation, and cell differentiation in plant growth? What are the different types of plant tissue? 3. What are the basic types of plant cells? 4. How do primary and secondary plant growth compare?