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The Cytoskeleton as a Regulator and Target of
The Cytoskeleton as a Regulator and Target of

... factors, and in many cases the plant cytoskeleton is instrumental in mediating the plant’s response. Cytoskeletal elements, for example, translocate chloroplasts under high light conditions (Takagi, 2000), facilitate gravity sensing (Blancaflor, 2002), and direct cellular response to wounding (Foiss ...
The syndapin protein family: linking membrane trafficking with the
The syndapin protein family: linking membrane trafficking with the

... protein that plays a crucial role in the uncoating of clathrincoated vesicles; Cremona et al., 1999) and synapsin I (a protein associated with the reserve pool of synaptic vesicles) (Hilfiker et al., 1999) (Fig. 1). These interactions first raised the possibility that syndapins have roles in both me ...
RICKETTSIA, CHLAMYDIA, MYCOPLASMA
RICKETTSIA, CHLAMYDIA, MYCOPLASMA

... life cycle, although they are also much smaller than bacteria. The first organism of this type was associated with pleuropneumonia of cattle, and was originally called the pleuropneumonia organism (PPO). Since that time, a number of organisms with similar morphological characteristics and cultural p ...
Ergastic
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Microtubules and the Evolution of Mitosis
Microtubules and the Evolution of Mitosis

... 2 Botanisches Institut 1, Kaiserstr. 2, 76128 Karlsruhe, Germany [email protected] ...
Prentice Hall Biology
Prentice Hall Biology

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Chapter 3C Notes from Book
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The plant cytoskeleton - The Company of Biologists
The plant cytoskeleton - The Company of Biologists

... Cells may expand in different ways: tip-growing cells, like root hairs, are usually exposed to the environment and are thought to expand only at the apical dome, whereas cells within tissues undergo a more uniform intercalary growth. The cyto­ skeleton may be specialized in different ways in support ...
to the complete text
to the complete text

... study the actin cytoskeleton Although yeast is not motile like Dictyostelium or higher eukaryotic cells, its our advanced knowledge of its genetics make it a good model system in which to study many aspects of the actin cytoskeleton. Cell division, secretion and signaling are well studied in the yea ...
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3D Visualization of Thylakoid Membrane
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Morphologic Investigation of Thunderstorm Initiates and GIS
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Questions on the integrity of the neuromuscular junction
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... locomotion and cellular integrity. Thus it appears that apoptosis is not a major factor in the senescent decline of C. elegans. Consistent with this conclusion, mutants defective in apoptosis do not exhibit lifespan changes (Hengartner, Exp. Geront., v.32, pp.363-374, 1997), and apoptotic gene expr ...
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... perform crucial functions. Plants exploit hydrodynamics to drive processes including gas exchange, leaf positioning, nutrient acquisition and growth. This paradigm is extended by recent work that reveals an important role for hydrodynamics in pollen tube growth. The pivotal role of water in the orig ...
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Cytoplasmic streaming



Cytoplasmic streaming, also called protoplasmic streaming and cyclosis, is the directed flow of cytosol (the liquid component of the cytoplasm) and organelles around large fungal and plant cells through the mediation of actin. This movement aids in the delivery of organelles, nutrients, metabolites, genetic information, and other materials to all parts of the cell. Cytoplasmic streaming occurs along actin filaments in the cytoskeleton of the cell.Cytoplasmic streaming was first discovered in the 1830s. The scientific breakthrough assisted scientists in developing an understanding of the different roles of cells and how they function as the basic operating systems of life.This process occurs through the operation of motor proteins called myosins.These proteins use energy of adenosine triphosphate (ATP) to act as a molecular motor, which slides along actin filaments. This works in a manner that tows the organelles and other cytoplasmic contents in the same direction. Myosin proteins consist of two conjoined proteins. If one protein remains attached to the substrate, the substance acted upon by the protein, such as a microfilament, has the ability to move organelles through the cytoplasm.The green alga genus Chara and other genera in the Division Charophyta, such as Coleochaete, are thought to be the closest relatives of land plants. These haploid organisms contain some of the largest plant cells on earth, a single cell of which can reach up to 10 cm in length. The large size of these cells demands an efficient means to distribute resources, which is enabled via cytoplasmic streaming.Cytoplasmic streaming is strongly dependent upon intracellular pH and temperature. It has been observed that the effect of temperature on cytoplasmic streaming created linear variance and dependence at different high temperatures in comparison to low temperatures. This process is complicated, with temperature alterations in the system increasing its efficiency, with other factors such as the transport of ions across the membrane being simultaneously affected. This is due to cells homeostasis depending upon active transport which may be affected at some critical temperatures.In plant cells, chloroplasts may be moved around with the stream, possibly to a position of optimum light absorption for photosynthesis. The rate of motion is usually affected by light exposure, temperature, and pH levels.In reference to pH, because actin and myosin are both proteins, strong dependence on pH is expected. The optimal pH at which cytoplasmic streaming is highest, is achieved at neutral pH and decreases at both low and high pH.The flow of cytoplasm may be stopped by:Adding Lugol's iodine solutionAdding Cytochalasin D (dissolved in dimethyl sulfoxide)↑ ↑ ↑ ↑ ↑ ↑
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