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Plant Tropism and Defenses • Lights important affect on plants – Photosynthesis… conversion of CO2 & H2O to energy containing carbon compounds and O2 – Phototropism… movement of plants toward light – Photomorphogenesis… the induction of important key development and growth stages due to the presence, intensity, and duration of light Light Frequencies • Action spectrum… the wavelengths useful to the plant – Red and blue light for most plants • blue-light photoreceptors – Active in phototropism & slowing hypocotyl elongation when seedlings break ground • Phytochromes – Activate de-etiolation process – Signals germination in many seed types depending on type of red light present (near vs far) » Allows plants to alter metabolism at different times of the day (shade avoidance) • Circadian clocks (rhythms) or biological cycles that are independent of extraneous environmental signals – Are internal to all Eukaryotic organsims • Can be reset by outside stimulus… light (Pr D Pfr) • Duration of night in some flowering plants – May be due to transcription factors that inhibit further production through allosteric inhibition. • Causes an increase in the morning and a fall in the evening – Clock will operate in the absence of day night although timing may be off Circadian Clock Response to External Stimuli • Gravitropism… causes stem to travel up and root to travel down – Orchestrated through statoliths (heavy starch grains) that settle in the bottom of cells • Causes a concentration of Ca2+ and auxin increasing growth on the upper portion resulting in bending of root • Thigmomorphogenisis (Mechanical stimuli)… – causes tree trunks to thicken and growth to be stunted – Plant leaves to curl when touched… action potentials resembling nerve impulses – Vines to grasp… thigmotropism Environmental Stresses • Drought… closes stomata and inhibits shallow root growth (deep root growth continues) • Flooding… fills pore spaces and drowns roots – Oxygen in the air spaces of soil is needed to support cellular respiration by root systems • Salt stress… reduces water potential of soil (becomes hypertonic to roots) dehydrates plant and reduces the plants ability to bring water to the stems and leaves – Some plants can produce solutes to counteract increases in salinity • Heat stress… denatures needed enzymes – Plants common to hot regions produce heat-shock proteins (chaperonins) to reduce the effect of the heat • Cold stress… affects solute transport as membranes become rigid – Production of unsaturated lipids increases (like cholesterol) to reduce crystallization and keep the membrane fluid Plant Defenses • Herbivory – Thorns – Toxins • kill of disable insect or animal • Attract predators that feed on possible eaters • Released to start production of toxins in neighboring plants