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29.1 Plant Nutrients
and Availability in Soil

Nutrients
 Elements
or molecules
essential for an organism’s
growth and survival

Plants require sixteen
elemental nutrients
available from soil, water,
and air
 Nine
macronutrients,
required in large amounts
 Seven micronutrients,
required in trace amounts
29.2 How Do Roots
Absorb Water and Nutrients?

Mycorrhizae = Mutualism between root and fungi in
which both species benefit



Root hairs = Thin extensions of root epidermal cells
that increase surface area for absorbing water and
mineral ions


Fungal hyphae share minerals absorbed from soil
Root cells provide fungus with food
New root hairs constantly form just behind the root tip
Root nodules = Masses of root cells infected with
bacteria that fix atmospheric nitrogen (N2) into a
form usable by plants (nitrogen fixation) NH3,
NH4+ and NO3-
How Roots Control Water Uptake



Osmosis drives water from soil
into the walls of parenchyma
cells of the root cortex
Water enters cell cytoplasm by
diffusion or through aquaporins;
active transporters pump
dissolved mineral ions into cells
Water and ions move from cell
to cell through plasmodesmata
The Casparian Strip


Endodermis between the cortex
and vascular cylinder secretes a
waxy substance which forms a
waterproof band (Casparian
strip) between plasma
membranes of endodermal cells
The Casparian strip forces
water and ions to enter the
vascular cylinder through
plasmodesmata or through
endodermal cell membranes
(controlled by transport
proteins)
Tracheids and Vessel Members
Water transport in plants

Transpiration = water loss from a plant through its leaves
Water leaves through open stomata
 Dry cell walls pull water from deeper in the leaf and pulls
water from the roots, stem and leaf through xylem (creates
negative pressure
 Water “sticks” to water (cohesion) and other molecules
(adhesion)
 Even small trees can lose 100L of water/day


What conditions increase transpiration???
Hot
 Dry
 Wind

Cohesion-Tension Theory
Controlling Water Loss at Stomata

Cuticle = A translucent, water-impermeable layer coating
the walls of all plant cells exposed to air



Consists of epidermal cell secretions: waxes, pectin, and cellulose
fibers embedded in cutin
Stomata = Openings through the plant epidermis that
regulate water vapor loss and gas exchange
Guard cells open or close the stoma depending on the
amount of water in their cytoplasm


Swollen cells open stoma
Collapsed cells close stoma
Homeostasis and stomata



Plants keep stomata open just enough so that gas
exchange can occur for photosynthesis but not so
much that they lose too much water
When water is abundant water flows into the leaf.
This increases water pressure in the guard cells and
opens them.
When water is scarce, pressure decreases and the
stomata close
Controlling Water Loss at Stomata

Environmental cues open or close stomata
 Water
availability (abscisic acid released by root cells
binds to guard cells and solutes and water leave)
 Carbon dioxide levels in leaf (aerobic respiration)
 Light intensity (triggers potassium pumps that pump
potassium into guard cells)
 Air pollution (prevents photosynthesis)