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What is evolution? Evolution • The relative change in the characteristics of a population over successive generations – Changes in traits etc. As time goes by. • Ruffled grouse has changed to become well camouflaged enabling it to survive • A population is the smallest unit that can evolve • Any shift in a gene pool is called evolution If evolution is the change in characteristics of a population over successive generations, then how do these characteristics change? • With the help of Adaptations and Variations Some Causes of evolution • 1. Adaptation – A particular structure, physiology or behavior that helps an organism survive and reproduce in a particular environment • Ex. Camouflage of tiger, excellent sense of smell, hearing, vision, etc • 2. Variation – A significant deviation from the normal biological form, function or structure • Ex. Albino moose, striped zebra.etc. Peppered Moth – Case of adaption (p.644) “Industrial Melanism • Peppered moth cones in two variations – Black( wing color is black – Flecked moths( light white wings with flecks of black) • Pre-mid 1900’s many more flecked moths than lack moths – Flecked moths would rest on lichens that provided camouflage. Black moths were easily seen and eaten • Industrial Revolution 'factories in England started producing black smoke that covered trees and killed lichens – Flecked moths were seen and eaten – Black moths survived long enough to reproduce and pass on their genes – Many more black moths than flecked moths. Frequency of genes has changed in the population. • 1950’s – England introduced the Clean Air policy ‘, less soot, lichens began to grow on trees again – 1959- 9/10 moths are black – 1985 5/10 moths are black – 1989 3/10 are black – 2010 black peppered moth will be as rare as before industrial revolution • How do peppered moths support evolution? – 1. 1850’s moths in England were mostly flecked (some were black) flecked ones survived because they were camouflaged – 2. 1900’s air pollution made trees black. Black moths were able to survive – 3. 1950’s saw pollution controls and now the flecked moth is surviving again Time line of evolution! • Evolution has taken a loooog time to occur. If 24hrs. Represented the entire evolutionary time scale. Humans would show up about 3 seconds before midnight • How long has it taken for evolution to occur and from what did it start? • Life is believed to have begun over 3500 million years ago with the earliest of cells. Mechanisms of evolution • Natural Selection – Proposed by Darwin – Idea where characteristics of a population change because individuals with certain heritable traits survive local environmental conditions and pass their traits onto their offspring – The environment determines which individuals are most fit to survive and pass on genes – Fitness- how well an organism fits its environment – Ex. Peppered moth, leaf bug Artificial Selection • Human selection of particular traits by breeding – Ex. Faster horses, disease resistant plants, dogs, etc • Humans determine the traits to be passed on to future generations • Population evolves in the direction man wishes it to • Not all breeding( artificial selection) is good. Breeding Pekinese and British bulldogs for flat face produces respiratory problems History of evolution. Who gave us what? • While we generally accept Darwin's theory of Natural selection as the mechanism of evolution several people have had an influence in evolutionary thought. 1. Greek philosophers • Greek philosophers such as Aristotle and Plato did not believe in evolution. They said all organisms which could exist were already created Georges Cuvier (1769-1832) • Founder of Paleontology(the study of fossils) • Fossil record revealed that something was causing species to appear and disappear • Thought that boundaries between fossils layers corresponded to catastrophic events such as Noah’s flood or droughts • Developed the theory of catastrophism • Catastrophes account for the disappearance and appearance of new species in the fossil record Charles Lyell (1797-1875) • Expanded on Hutton's idea of Gradualism • Gradualism– idea that earth’s geological features are in a slow continual cycle of change • Developed the theory of Uniformitarianism – Idea that geological processes operated at the same rates in the past as they do today. • He rejected the idea of catastrophism, etc • He said the world was millions of years old and not 6000 years as believed Thomas Malthus (1789) • Looked at plants and animals • Said that plants and animals grow faster than their food supply • Causes a population to be reduced by starvation, disease, etc. • Crowding and struggle for food and resources is what kept population from exploding • Darwin borrowed ideas on struggle for survival and realised those with the best traits for survival would pass on their gene Jean Baptiste Lamarck (1744-1829) • Published a theory of evolution in 1809 the year Darwin was born • Believed that organisms came from nonliving sources • Said that organisms respond to the needs in their environment • Proposes the idea that body parts used extensively to cope in the environment would be come stronger and stronger (idea of use and disuse) – Examples biceps of blacksmiths ,giraffes neck • He based his theory on two observations thought to be true in his day – 1. use and disuse• individuals lose characteristics they do not require and develop those which are useful. – Examples such as the black smith biceps – 2. Inheritance of acquired traits • Individuals inherit the acquired traits of their ancestors – Ex. A child would be strong because their dad was a weightlifter – A person who accidentally lost a finger would produce offspring with nine fingers Alfred Wallace (1858) • British Naturalist developed same theory as Darwin • Wallace did extensive fieldwork, first in the Amazon River and the Malay Archipelago, where he identified the Wallace Line that divides the Indoneaian archipelago into two distinct parts: a western portion in which the animals are largely of Asian origin, and an eastern portion where the fauna reflect Australasia • He was considered the 19th century's leading expert on the geographical distribution of animal species and is sometimes called the "father of biogeography“.] • Wallace was one of the leading evolutionary thinkers of the 19th century and made many other contributions to the development of evolutionary theory besides being co-discoverer of natural selection. These included the concept of warning coloration in animals, and the Wallace effect, a hypothesis on how natural selection could contribute to speciation by encouraging the development of barriers against hybridization. Charles Darwin • In 1831 he left on a 5 year voyage on board the beagle • He stopped in the Galapagos islands where the diversity of tortoises and birds amazed him • His theory of Descent with Modification had two main ideas – 1. present life forms have risen by descent and modification from an ancestral species – 2. Natural selection is the mechanism of modification over long periods of time • He returned to England in 1836 • He wrote the book `the origin of the Species` in which he published his theory of evolution Natural Selection and Evolution • • • • Summary of Darwin's Findings on Galapagos Islands Noticed that each island had finch birds that were different from each other Noticed that tortoises were different on each of the islands If these were the products of creation, how could such variations have occurred in such a small area Darwin thought that these organisms must have evolved from a common ancestor Darwins Finches Darwin's Theory of Natural Selection • Main points – 1. organisms produce more offspring than can survive – 2. competition occurs between individuals for limited resources. This causes a struggle to survive – 3. There are variations in individuals in a given population and these traits can be passed on. Variations are often caused by Mutations – 4. only the individuals that are better suited to local environmental conditions survive to reproduce What Darwin could not Explain • Darwin was not able to explain how the favorable traits were passed on to the offspring • why – He knew nothing of Mendel (heredity) and Mutations(producing variations) that could be passed on – However Mendel's ideas supported Darwin's ideas. This produced a revised theory of evolution Theory of Modern Evolution or Modern synthesis • This is the theory of evolution commonly accepted today. • 1. This is a meshing of Mendel's and Darwin's ideas • 2. Darwin says that variations exist in a population allowing certain organisms to adapt to their environment. These traits can then be passed on to offspring • 3. Mendel's work points out that mutations are the cause of variation within a population and it is the DNA that helped carry these best traits onto the next generation Evidence Supporting the Modern Theory of evolution • The following are pieces of evidence that supports the modern theory of evolution – 1. Fossil record – 2. Biogeography – 3. Comparative Anatomy • A. Homologous structures • B. Analogous structures • C. Vestigial structures – 4. comparative embryology – 5. heredity – 6. Molecular biology 1. Fossil Record Fossil: remains or traces of once living organism. Often preserved in rock. • Fossil evidence supports evolution in the following ways – A. Fossils from more recent geological eras are more similar to present day organism than older fossils • This supports the idea that life evolve over time – B. Fossils appear in chronological order in sedimentary rock. Younger fossils appear higher in sedimentary layers and are more complex than older fossils appearing in deeper layers • This supports the idea that has evolution has occurred , species of organisms have become more complex – C. Transitional Fossils make links between different sets of related organisms within differing fossil layers • Ex. Archaeopteryx shows a relationship between reptiles and birds. • This suggests that evolution is occurring over time from less complex to more complex life forms. Finding the Age of Fossils Dating Fossils • There are two methods to determine the age of fossil. • A. Relative Dating; Judging the age of a fossil according to its position in the layer of rock. – Ex. Fossil B is younger than C but older than Fossil A A B C • B. Absolute Dating-finding the exact age of a fossil using radioactive dating – Radioactive dating- a method of finding the age of a fossil using half life of certain radioactive substances that decay over time – Note: The radioactive substances decays into a more stable daughter element – Half Life: period of time required for 1./2 of a radioactive isotope to decay into a more stable element • Representative radioactive isotopes with half life Radioactive Parent Stable daughter Half Life (years) C14 N14 5730 U235 Pb 207 713000000 K40 Ar 40 1250000000 Rb 87 Sr 87 48800000000 C 14 C14 N14 N14 C14 C14 5730 years 5730 years One ½ life One ½ life Notice after each half life only ½ the original C14 sample remains. The other ½ has been changed into N14( the more table form Percentage of original sample remaining 100% 50% 1 half life 1 25% 2 half life 1/2 12.5% 6.25% 3.125% 3 half life 1/4 4 half life 1/8 5 half life 1/16 1/32 1.56% 6 half life 1/64 Calculations involving half life • A. Finding amount of sample remaining • Procedure: use ½ n (where n=# half flies) to find multiplication factor. Then multiply total by original mass – Ex. A 10 kg sample of C14 has underwent 4 half life's. How much of the original sample remains? • Ans. ½ n = ½ 4 = ½ x ½ x ½ x 1/2 = 1/16 • Now multiply 1/16 x original mass • 1/16 x 10kg= .625 kg remaining Finding the half life( time required for a substance to deacy1/2 its original amount • Ex. A rock is found to be 33000000 years old and contains 1/64 of the original sample. What is the half life of the rock/ • First find the number of half life's it took to reduce the sample to 1/64 • To do this we ask the question, ½ to what power =1/64/ the easiest way to find this is to ask the question , 2 to what power is 64. In this case 26 =2x2x2x2x2x2, so we conclude that the number of half life's =6 • Next divide the age of the rock by the number of half life's and you will find the value of 1 half life • In this case 33000000/6 half-life's = 5500000/ half life's Finding the age of the fossil • Ex. A fossil contains 1/32 of the original U-235. what is the age of the fossil if the half life of u-235 is 713000000? • Answer – First we need the number of half life's that reduces the fossil to 1/32 of its original amount – To do this we as the question ½ to what power is 1/32. Obviously it would be ½ to the power of 5. This means that 5 half life's have passed. • Next we use the 5 half life's and multiply the value of a single half life to get fossil age. In this case we have the following • 5 half lifes X 713000000yrs/ half life= 3565000000 yrs old 2. Biogeography • This is the study of the geographical distribution of species • Darwin noticed that the birds on the Galapagos islands were similar to those on the mainland of South America • Geographically close environments (desert and jungles of south America) are more likely to be populated by related species rather than locations that are geographically separate but environmentally similar ( desert of Australia and a desert in Africa) 3. Comparative Anatomy • This is a comparison of physical structures in differing organisms that may suggest a common ancestor. These methods are looked at: • Homologous Structures: these are body structures in different species which have the same origin but differ in structure and function. – Ex. Human arm, frog leg, bat wing, horse leg • These structures all have a similar number of bones/ ligaments suggesting they came from a common ancestor, but they all have a different structure and function • Analogous structures: these structures that have different origins but similar functions – Ex. Bird and incest wings • Vestigial structures: these are structures that were functional in ancestors, but have no current function – Ex. Pelvic bone in baleen whales, wings in ostriches, appendix in humans Homologous structures Analogous structures Vestigial 4. Comparative Embryology • This is a comparison of embryos from various species to indicate relationships among organisms • Many embryos have similar stages of development – Ex. All vertebrates go through a stage having a gill pouch Comparative embryology 5. Heredity • Knowledge of heredity can explain how variations can occur in a population allowing members of that population to be better suited to their environment and thus undergo natural selection 6. Molecular Biology • This is a comparison of the DNA and proteins within various species to indicate relationships/ similarities • The closer the DNA sequences are between organism the more closely related the species are. This may suggest a common ancestor • Ex. Humans and chimpanzee differ by only 2.5% Population Genetics and Hardy Weinberg • Population genetics – This is a study of the genes in a population and how they may or may not change over time • Recall: if there is a shift in the gene pool of a population them we know evolution is happening • Population – A localized group of a single species occupying a particular area • Gene pool – This is the total of all genes within a population Hardy-Weinberg Principle • Proposed by Hardy and Weinberg • A model of a population that is not changing to help understand a population that is changing • Premise of the principle – The principle states that in a population under certain conditions the frequency of alleles will remain stable from generation to generation • In other words, under certain conditions a populations genetic makeup will not change meaning it is not evolving. It is in Genetic Equilibrium • The principle explains why recessive alleles do not disappear in a population over time and to helps explain why dominant traits do not become more widespread Conditions necessary to establish a population hardy Weinberg equilibrium • • • • Requirements 1. No mutations occur in the population. 2. No immigration or emigration. 3. There must be a very large population in order to avoid genetic drift. • 4. There must be no natural selection No genotype has an advantage over another. • 5. There must be no sexual selection mating is random. Formula • 1. p+ q= 1 • P= frequency of dominant allele(how often the dominant allele shows up in the total population • q= frequency of the recessive allele(how often the recessive allele shows up in the total population • 1=100% • Since there are only ever two alleles for a trait , the total amount of the allele always has to add up to be 100% or 1 • P2 + 2pq +q2 = 1 • P2 = frequency of Homozygous dominant genotype • 2pq=frequency of heterozygous genotype • q2 frequency of homozygous recessive genotype example • Suppose we have a population of Gerbils with he following conditions Phenotype Black Black White Genotype BB Bb Bb Number of gerbils 196 168 36 Total number 400 of gerbils (192+168+36) = 400 400 400 Genotype frequency (BB,Bb,bb) Bb=168/400= 0.42 BB= 36/400=0.09 BB= 196/400=0.49 Allele frequency (B and b) B= 196 + 168/800=0.07 b= 168 + 36/800 = 0.03 • The table above shows the frequencies and genotypes frequencies . Notice how they are calculated • Now lets look at formula • we know BB = 0.49= p2 • Bb = 0.42= 2pq • Bb = 0.09=q2 • According to the formula • If we find want to find the frequencies of the alleles B and b we need to find the square root of p2 and q2 so More on Evolution • Microevolution - population change in allele frequencies • Macroevolution – grand scale changes as seen in the fossil record Mechanisms of evolution • The following are mechanisms that cause genetic variation in a population and thus move them away from Hardy – Weinberg equilibrium • In other words the following cause populations to evolve – – – – – – Mutations Genetic drift Gene flow Non-random mating Natural selection Sexual selection Mutations • Changes in the DNA that bring new alleles in a population • New alleles provide variations that cause evolution • Mutations can be harmful, neutral or beneficial • Mutations are beneficial if they provide a selective advantage which allows certain organisms to adapt to their environment • Ex. California ground squirrel having the ability to break down rattlesnake poison Genetic Drift • Change in allele frequencies in small populations caused by chance alone – For example in a small population mutations can cause allele frequencies to change whereas in a large population the mutations may have little to no effect on the frequencies. The gene pool will not shift if the population is large • The allele frequencies in small populations can change over time and this can lead to evolution. Remember: in a non evolving population ( hardy Weinberg equilibrium) the allele frequencies remain unchanged Causes of Genetic drift • A. Bottleneck effect: a situation in which as a result of chance some alleles are overrepresented and others are underrepresented because a population has been reduced through natural disasters etc. – Ex. Elephant seals have passed through a bottleneck. They have been overhunted causing their numbers to be reduced to about 20. because of this certain alleles have been eliminated)( variety reduced). The population has since grown to 30000 having little variation. This has resulted in a change in the allele frequency • Founder effect • When a small amount of organisms (called a founder population) move into a new area, chances are they do not contain the entire population in the parent population this results in a changes in allele frequencies – Ex. Hawaiian honey creeper birds migrated from north America Polydactyl in Quakers Gene Flow • This is the movement of genes into or out of a gene pool • This causes a change in the gene pool resulting in evolution • If gene flow happens enough between two neighbouring populations they may eventually merge into one population with a common genetic structure Non Random Mating • If a population mates on a random basis genetic equilibrium is maintained and the population does not evolve • Normal populations do not undergo random mating. For example individuals will mate more with their neighbours rather than distant organisms there are two types of non random mating • Inbreeding- mating between closely related organism – This will cause a loss in variety in the population and the allele frequencies will change • Assortive Mating- this is where organisms choose mates that are similar to themselves – Artificial selection ( breeding of certain dogs) is an example of assertive mating. The dogs being mated are choosing (or are chosen for them) mates that are similar to themselves – This causes a reduction in variety in the population causing allele frequencies to change Non-Random Mating Natural selection • A populations characteristics can change because certain individuals within the population have heritable traits that allow them top adapt to and survive local environmental conditions • There are 3 types of Natural Selection – Stabilizing selection – Directional selection – Disruptive selection Stabilizing • Natural selection where an intermediate or normal phenotype is favored over the extremes – Ex. Birth weight( most babies born today are of average or normal weight because the extreme (low or high) birth weight babies are selected against(we do not see many low birth weights or high birth weights babies anymore) The middle intermediate phenotype is favored Directional • Selection where one extreme phenotype is favored over the other. This causes a shift in the phenotypes in that direction • This type of selection is common during environmental change or when population migrates to a new habitat – Ex modern horse (adapted to a grassland habitat) adapted from an ancestral horse (adapted to a forest habitat). Most horse today resemble the modern horse and not the forest horse Disruptive • Selection where both extremes of the phenotype are selected rather than the middle (intermediate phenotype) • The intermediate phenotype may be eliminated from the population – Ex. Coho salmon. Males are either small (jack salmon) or very large. No real medium size male salmon found in the population Sexual Selection • This is selection on the basis off being able to find a suitable mate in which to produce offspring. Having the ability to choose a mate helps ensure genetic information is passed on as well as introduces variety into the population • Finding a suitable mate is based on 2 main characteristics – Male competition- male competition can determine who gets the chance to mate with female – Female choice-females choose who they mate with Sexual Selection • Sexual selection acts on an organism's ability to obtain or successfully copulate with a mate. • Selection makes many organisms go to extreme lengths for sex: Speciation • Formation of a species • Biological species: a group of organisms able to interbreed and produce fertile offspring – Ex. Horses and donkeys are separate species. They are able to interbreed, but the offspring produced are not fertile How do species form? There are generally two pathways in which species are formed • 1. Transformation- the formation of a species because of a series of accumulated changes over time. One species changes into another in this way. – Species changes new species • 2. Divergence- the formation of species from a parent species/ ancestor – Parent species » Species Speciation occurs when two groups become isolated from each other Types of Speciation • Allopatric—separation of members of the same species by a physical barrier. The separate populations over time may evolve distinctly different characteristics. If the geographical barriers are later removed, members of the two populations may be unable to successfully mate with each other, at which point, the genetically isolated groups have emerged as different species. • Allopatric isolation is a key factor in speciation and a common process by which new species arise.[ • Parapatric—Occurs in adjacent populations due to local environment problems. is the relationship between organisms whose ranges do not significantly overlap but are immediately adjacent to each other; they only occur together in a narrow contact zone. • Sympatric—is the process through which new species evolve from a single ancestral species while inhabiting the same geographic region • individuals continue to live with each other. Mostly in plants. Due to polyploidy. Allopartric vs. Sympatric What causes species to become isolated? Barriers • Types of barriers – Geographical Barriers- when a population becomes divided by a geographical boundary such as a canyon, river, etc. This prevents interbreeding. Over time natural selection causes genetic differences to become so large two species form. • Ex. Giraffes that become separated by mountains will eventually develop into separate species Biological Barriers • These are barriers that keep species reproductively isolated • These barriers may be Pre-zygotic barriers or Post zygotic Barriers Pre zygotic Barriers These are known as pre-fertilization barriers that either impede mating or prevent fertilizations • a. Behavioural isolation- bird songs, courtship rituals pheromones, etc. Are all species specific and prevent fertilization • b. Temporal isolation; these are usually timing barriers. Several species mate at different times during the year and as such are not able to mate • c. Habitat isolation-some species live in the same area but have different habitats – Ex. North American garter snakes. One prefers open areas while the other prefers water • d. Mechanical isolation- some species are automatically incompatible thus not allowing them to exchange sperm and egg – Ex. The genitals on certain species on insects work on a lock and key hypothesis. If the lock does not fit the key, no fertilization can happen • e. Gametic Isolation-sometimes the gametes from species do not even meet. This prevents fertilisation. – Ex. Sea urchins release eggs into the water but chemicals on the surface of the eggs prevent sperm from as different species to fertile them. Post Zygotic Barriers These are barriers that prevent a zygote from developing into a fertile organism • a. Hybrid Invariability-incompatibility of two species may cause the zygote to stop embryonic development. – Ex. Embryos of sheep/goats do not survive • b. Hybrid sterility- this is the production of an organism but it is sterile – Ex. Horse + donkey=mule(sterile) • c. Hybrid Breakdown-sometimes the first generation of offspring are viable and can reproduce, but when their offspring reproduce their offspring are sterile or weak. – Ex. Cotton plants produce generations of seeds that die Patterns of Evolution • Convergent Evolution—become more alike due to environment. Aquatic mammals and fish. • Divergent Evolution—Share a common ancestor, but evolve differently. • Coevolution—Plants and pollinators; parasites and hosts. Adaptive Radiation • The diversifying of an ancestral species into a variety of species • This usually occurs after a novel characteristic has evolved or if there is a mass extinction Species 1 Species 2 • Ancestral species Species 3 – Ex. Galapagos finches from one common ancestor on islands Adaptive Radiation • The finches of the Galapagos Islands provide a classic example this evolutionary process - a single lineage gives rise to species occupying diverse environmental niches. (13 species) Divergent evolution • This is where species that were once similar to an ancestral species diverge to become different species • Note: Adaptive radiation is an example of Divergent Evolution Convergent evolution • This is evolution where two completely unrelated species have similar traits – Ex. Birds and bees have wings (similar trait) • Each species develops the same traits because they adapt to the same type of environmental conditions • The species do not come from a common ancestor Co evolution • This is evolution where two species change together where each species responds to changes in the other – Ex. Milkweed plants and monarch butterflies. The milkweed plant has toxins in their leaves. Monarch butterfly eats the leaves and absorbs the toxins making them toxic. Most birds avoid monarch butterflies for this purpose Pace of evolution • How fast does evolution occur? • There are two theories that explain how fast evolution occurs. Both examine the fossil record. Gradualism • A model that says change occurs slowly and steadily before and after a divergence • Fossils show a slow and repeated change through the fossil record Punctuated equilibrium • A model; proposed by Gould and Eldridge • A model that proposes evolution happens in spurts • The model says that species undergo long periods of stasis where they remain unchanged followed by short periods of very rapid change (spurts) • The changes are usually brought about by sudden environmental changes such as volcanoes earthquakes etc. • Species previously disadvantaged could now be advantaged and new species could develop quickly Origins of the World and Life • Many theories exist that try to explain the origin and development of life on earth • The following will be considered – – – – – – Chemical evolution’ Panspermia Gaia hypothesis Heterotroph hypothesis Symbiogensis (symbiotic theory) Intelligent design Chemical Evolution • A theory of evolution created by Oparin-Haldane • They said that organic molecules (the building blocks of life) could develop from inorganic compounds present on the surface of the early earth – The earth had an atmosphere that consisted of no oxygen, but plenty of hydrogen, Ammonia(NH3 , Methane and water vapour(inorganic molecules) – These gases condensed and formed a primordial soup – Energy from, lightening and UV radiation caused organic molecules to develop from inorganic molecules in the soup – Overtime the organic molecules combined to become an early life form • Miller Urey • Two scientists who designed an experiment to prove Oparin-Haldanes theory. Here is what they did • They combined methane, ammonia, water vapour and hydrogen in a flask and exposed the gases to an energy source simulating lightening • The liquid inside the flask changed color and when examined contained several organic compounds including amino acids – Amino acids make up protein which make up the structure of most living things Panspermia • A theory that suggests life began elsewhere in the universe and migrated to our plant • For example it is believed that life originated from bacterial cells elsewhere and travelled from outer space to earth on meteorites Gaia Hypothesis • theory put forward by James Lovelock • Idea that earth is a super organism called Gaia • The earth has systems that keep a balance between temperature and atmosphere • After life originated on earth, Gaia came alive and began to regulate earth systems • The systems help provide an environment where life could exist and survive Heterotroph Hypothesis • Theory put forth by Oparin – Said that firs cells on earth had to be heterotrophs that eventually developed into autotrophs • Primordial soup existed of organic molecules • The environment was oxygen poor • Heterotrophs such as anaerobic bacteria fed on the organic molecules • The hetrotrophs began to release carbon dioxide into the atmosphere, The heterotrophs developed into autotrophs and began using carbon dioxide • The autotrophs began to release oxygen into the atmosphere • This made the atmosphere oxygen rich that could now support life Symbiogensis • Put forth by Lynn Margulis – Theory that attempts to explain the development of mitochondria and chloroplasts as organelles that appear in eukaryotic cells – Chloroplasts and mitochondria have their own DNA and come from the symbosis (working together) of prokaryotic cells – Here is how she said eukaryotic organisms developed • An anaerobic bacterium ate but did not digest an aerobic bacterium(called a guest bacterium) • The guest bacterium provided oxygen to the bacterium. The guest bacterium eventually became a mitochondrion • Other bacteria ate photosynthesizing bacteria. The photosynthesizing bacteria became chloroplasts Intelligent Design • Theory suggests that life and mechanism of life are too complex to have evolved by chance • Believed that the generation and evolution of life must have been directed by some unidentified supernatural intelligence