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2. Increased metabolic heat production occurs when a. hormonal changes boost the metabolic rate in birds and mammals, b. birds and mammals shiver, c. organisms increase their physical activity, and d. honeybees cluster and shiver. 3. Insulation is provided by a. hair, b. feathers, and c. fat layers. 4. Circulatory adaptations include a. increased or decreased blood flow to skin and b. countercurrent heat exchange, with warm and cold blood flowing in opposite directions. 5. Evaporative cooling may involve a. sweating, b. panting, or c. spreading saliva on body surfaces. 6. Behavioral responses a. are used by endotherms and ectotherms and b. include i. moving to the sun or shade, ii. migrating, and iii. bathing. 25 Control of Body Temperature and Water Balance CHAPTER Lecture Outline I. Introduction (Bears Essay) A. During cold winters, bears are often dormant. B. Physiological processes aid homeostasis, keeping the body temperature about 5ºC below normal. 1. Body fat and dense fur provide insulation. 2. Blood flow to extremities is reduced. 3. Nitrogen-containing wastes are metabolized differently. C. Homeostasis is the maintenance of steady internal conditions despite fluctuations in the external environment. D. Examples of homeostasis include 1. thermoregulation—the maintenance of internal temperature within narrow limits, 2. osmoregulation—the control of the gain and loss of water and solutes, and 3. excretion—the disposal of nitrogencontaining wastes. II. Thermoregulation A. 25.1 An animal’s regulation of body temperature helps maintain homeostasis 1. Thermoregulation is a. the process by which animals maintain an internal temperature within a tolerable range and b. a form of homeostasis. 2. Ectothermic animals a. gain most of their heat from external sources and b. include many fish, most amphibians, lizards, and most invertebrates. 3. Endothermic animals a. derive body heat mainly from their metabolism and b. include birds, mammals, a few reptiles and fish, and many insects. B. 25.2 Heat is gained or lost in four ways 1. Heat exchange with the environment may occur by a. conduction—the transfer of heat by direct contact, b. convection—the transfer of heat by movement of air or liquid past a surface, c. radiation—the emission of electromagnetic waves, or d. evaporation—the loss of heat from the surface of a liquid that is losing some of its molecules as a gas. C. 25.3 Thermoregulation involves adaptations that balance heat gain and loss 1. Five general categories of adaptations help animals thermoregulate. III. Osmoregulation and Excretion A. 25.4 Animals balance the level of water and solutes through osmoregulation 1. Osmoregulation is the homeostatic control of the uptake and loss of water and solutes such as salt and other ions. 2. Osmosis is one process whereby animals regulate their uptake and loss of fluids. 3. Osmoconformers a. have body fluids with a solute concentration equal to that of seawater, b. face no substantial challenges in water balance, and c. include many marine invertebrates. 4. Osmoregulators a. have body fluids whose solute concentrations differ from that of their environment, b. must actively regulate water movement, and c. include i. many land animals, ii. freshwater animals such as trout, and iii. marine vertebrates such as sharks. 5. Freshwater fish a. gain water by osmosis (mainly through gills), b. lose salt by diffusion to the more dilute environment, c. take in salt through their gills and in food, and d. excrete excess water in dilute urine. 6. Saltwater fish a. lose water by osmosis from the gills and body surface, b. drink seawater, and c. use their gills and kidneys to excrete excess salt. 7. Land animals a. face the risk of dehydration, b. lose water by evaporation and waste disposal, c. gain water by drinking and eating, and d. conserve water by i. reproductive adaptations, ii. behavior adaptations, iii. waterproof skin, and iv. efficient kidneys. B. 25.5 EVOLUTION CONNECTION: A variety of ways to dispose of nitrogenous wastes has evolved in animals 1. Metabolism produces toxic by-products. 2. Nitrogenous wastes are toxic breakdown products of proteins and nucleic acids. 3. Animals dispose of nitrogenous wastes in different ways. 4. Ammonia (NH3) is a. poisonous, b. too toxic to be stored in the body, c. soluble in water, and d. easily disposed of by aquatic animals. 5. Urea is a. produced in the vertebrate liver by combining ammonia and carbon dioxide, b. less toxic, c. easier to store, and d. highly soluble in water. 6. Uric acid is a. excreted by some land animals (insects, land snails, and many reptiles), b. relatively nontoxic, c. largely insoluble in water, d. excreted as a semisolid paste, conserving water, but e. more energy expensive to produce. C. 25.6 The urinary system plays several major roles in homeostasis 1. The urinary system a. forms and excretes urine and b. regulates water and solutes in body fluids. 2. In humans, the kidneys are the main processing centers of the urinary system. 3. Nephrons a. are the functional units of the kidneys, b. extract a fluid filtrate from the blood, and c. refine the filtrate to produce urine. 4. Urine is a. drained from the kidneys by ureters, b. stored in the urinary bladder, and c. expelled through the urethra. D. 25.7 Overview: The key processes of the urinary system are filtration, reabsorption, secretion, and excretion. 1. Filtration a. Blood pressure forces water and many small molecules through a capillary wall into the start of the kidney tubule. 2. Reabsorption a. refines the filtrate, b. reclaims valuable solutes (such as glucose, salt, and amino acids) from the filtrate, and c. returns these to the blood. 3. Substances in the blood are transported into the filtrate by the process of secretion. 4. By excretion the final product, urine, is excreted via the ureters, urinary bladder, and urethra. E. 25.8 Blood filtrate is refined to urine through reabsorption and secretion 1. Reabsorption in the proximal and distal tubules removes a. nutrients, b. salt, and c. water. 2. pH is regulated by a. reabsorption of HCO3– and b. secretion of H+. 3. High NaCl concentration in the medulla promotes reabsorption of water. F. 25.9 Hormones regulate the urinary system 1. Antidiuretic hormone (ADH) regulates the amount of water excreted by the kidneys by a. signaling nephrons to reabsorb water from the filtrate, returning it to the blood, and b. decreasing the amount of water excreted. 2. Diuretics a. inhibit the release of ADH and b. include alcohol and caffeine. G. 25.10 CONNECTION: Kidney dialysis can be lifesaving 1. Kidney failure can result from a. hypertension, b. diabetes, and c. prolonged use of common drugs, including alcohol. 2. A dialysis machine a. removes wastes from the blood and b. maintains its solute concentration. Vocabulary to Know ammonia filtrate renal medulla antidiuretic hormone filtration secretion (ADH) glomerulus (plural, glomeruli) thermoregulation Bowman’s capsule loop of Henle urea collecting duct nephron ureter countercurrent heat exchange osmoconformer urethra dialysis osmoregulation uric acid distal tubule osmoregulator urinary bladder ectotherm proximal tubule urinary system endotherm reabsorption urine excretion renal cortex Word Roots anti- = against; -diure = urinate (antidiuretic hormone: a hormone made by the hypothalamus and secreted by the posterior pituitary that promotes water retention by the kidneys) counter- = opposite (countercurrent heat exchange: parallel blood vessels that convey warm and cold blood in opposite directions, maximizing heat transfer to the cold blood) dia- = through; -lyso = loosen (dialysis: the separation and disposal of metabolic wastes from the blood by mechanical means; an artificial method of performing the functions of the kidneys) ecto- = outside; -therm = heat (ectotherm: an animal that warms itself mainly by absorbing heat from its surroundings) endo- = within; -therm = heat (endotherm: an animal that derives most of its body heat from its own metabolism) glomer- = a ball (glomerulus: the ball of capillaries in the nephron of the vertebrate kidney surrounded by Bowman’s capsule; together, the glomerulus and Bowman’s capsule produce filtrate from blood) osmo- = pushing; -conform = the same (osmoconformer: an organism whose body fluids have a solute concentration equal to that of its surroundings); -regula = regular (osmoregulation: the control of the gain and loss of water and dissolved solutes in an organism; osmoregulator: an organism whose body fluids have a solute concentration different from that of its environment and that must use energy to control water loss or gain) reni- = a kidney; cortex- = shell (renal cortex: the outer portion of the vertebrate kidney; renal medulla: the inner portion of the vertebrate kidney) thermo- = heat; -regula = regular (thermoregulation: the maintenance of internal temperature within a range that allows cells to function efficiently) Name Date Class Ch. 25 Study Guide: On a separate sheet of paper- Use the Chapter 25 Outline to construct a summary for each of the following objectives by converting the appropriate information into paragraph form. Opening Essay Explain how bear physiology adjusts during a period of dormancy. Thermoregulation 25.1 Distinguish between endotherms and ectotherms, providing examples of each. 25.2 Describe the four ways that heat is gained or lost by an animal. 25.3 Describe the five general categories of adaptations that help animals thermoregulate. Provide specific examples of each. Osmoregulation and Excretion 25.4 Describe the osmoregulatory challenges and associated adaptations of freshwater and saltwater fish, terrestrial arthropods, and terrestrial vertebrates. 25.5 Describe the three ways that animals eliminate nitrogenous wastes, and the advantages and disadvantages of each method. 25.6 Describe the general and specific structure of the human kidney. Explain how this organ promotes homeostasis. 25.7 Describe the four major processes by which the human excretory system produces and disposes of urine. 25.8 Describe the key events in the process of converting filtrate into urine in the kidneys. 25.9 Explain how antidiuretic hormone contributes to homeostasis. 25.10 Explain how a dialysis machine functions.