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Phylum Annelida Latin for “Little rings” Segmented Worms Segmentation  Repeating, specialized sections for different purposes or functions  Coelom is divided into separate compartments by partitions  Septa  Enables different compartments to contract or expand independently  Duplication of some of the organ systems between each segment provides insurance against injury Taxonomy  Class Polychaeta   Class Hirudinea   many bristles leeches Class Oligochaeta  earthworms Class Polychaeta - “many bristles”    Mostly marine worms Have paddle-like appendages on sides  “parapodia”  Used for movement and respiration Some have bristles to aid in defense Bristleworm – many setae Class Polychaeta  Have antennae   Sense the environment Have specialized mouthparts for feeding  Jaws (free swimming predators) Class Polychaeta  Some free living    E.g. Nereis Predatory Some are sessile   live in tubes made of calcium, silica (sand) or protein Filter feeders Class Hirudinea   Smallest class of Annelids (~300 species) Leeches    Suck vertebrate blood Carnivorous Powerful suckers at both front and back ends Leeches  No setae or parapodia   Have suckers at anterior and posterior ends Move by “looping”:   Attach anterior sucker then pull rest of body forward Classified as a parasite    Secrete an anaesthetic that prevents the host from feeling their presence Secrete an anti-clotting agent If undisturbed, a leech can ingest 10 times its own weight in blood Class Oligochaeta  Earthworms    Found in soil and freshwater Some types are found in the ocean Tubifex (sludgeworm)  Can survive in polluted sediments, areas with little oxygen Oligochaeta anatomy Oligochaeta anatomy Movement  Requires coordinated movement between the circular and longitudinal muscles  To move, the worm:  anchors middle segments using setae  contracts circular muscles in front of the anchored segments  this “squeezes” the fluid in the coelom which in turn causes the worm to “stretch” forward  the worm then anchors the anterior end, releases the middle and uses the longitudinal muscles to pull forward the posterior end  Made possible by segmentation Movement Feeding and Digestion  Ingest soil as they burrow  Prostomium – flap of skin that covers the mouth when not feeding Mouth – where soil enters the worm Pharynx – muscular region that forces soil through esophagus into the crop    Crop – serves as temporary storage  Gizzard – muscular “blender” that grinds soil and organic matter Feeding and Digestion  Intestine – long “tube” that absorbs any nutrients in the ground-up soil   has a special fold called a Typhlosole  increases the surface area of the intestine so that nutrients can be absorbed more efficiently Anus – where any undigested material exits the worm = “worm manure” Feeding and Ecology  Earthworms are decomposers  Breakdown organic matter and  Release nutrients into soil , thus fertilize it  Loosen soil as they burrow and  Aerate it (increase its oxygen content) Feeding and Digestion prostomiumm Circulation  Closed circulatory system  transports oxygen, carbon dioxide, nutrients and wastes through the body of the earthworm  Ventral and Dorsal blood vessels  Blood is forced through blood vessel by contraction of vessels and 5 aortic arches Circulation Respiration  No specialized respiratory organs  Oxygen and Carbon dioxide diffuse directly through skin and into small blood vessels  Only if skin is moist Excretion  Nephridia or Nephrostome act like kidneys   remove waste and control water levels in blood Nephridiopores    pores on ventral side of worrm release waste and excess water each segment except first three and last one have nephridia and nephridiopores Excretion Nervous System  Chain of ganglia connected by a ventral nerve cord  Most body segments have a single ganglion  Have an anterior cerebral ganglia (brain!)  Can respond to light, touch, chemicals, moisture, temp, vibrations Nervous System Reproduction  Hermaphroditic  individual worm cannot fertilize own eggs Reproduction     The “Position”  head to tail, ventral surface to ventral surface The “Transfer”  clitellum secretes mucus  each earthworm injects sperm into the mucus  mucus moves into pouch-like seminal receptacle of each worm The “Delivery”  several days later the worm secretes a tube of mucus and chitin (thick carbohydrate) from clitellum  as the worm moves the tube “picks” up eggs and carries it to sperm to be fertilized The “Result”  tube closes up to form protective case and 2-3 weeks “baby” worms hatch Review