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Phylum Echinodermata Introduction Echinodermata • About 6000 species, all marine Echinodermata Major characteristics • secondary pentamerous radial symmetry internal skeleton • water vascular system Water Vascular System • • • • • • • Madreporite stone canal ring canal radial canal lateral canals Ampulae tube feet CLASSIFICATION OF ECHINODERMATA • • • • • Class Asteroidea Class Ophiuroidea Class Echinoidea Class Holothuroidea Class Crinoidea Class Asteroidea True Starfishes Class Asteroidea • arms not sharply delineated from central disc • tube feet with suckers; used for – Locomotion – obtaining food • madreporite and anus aborally located • some have pedicellariae jawlike appendages of epidermis True Starfishes Class Asteroidea True Starfishes • Feeding – Mouth – cardiac stomachcan be extruded – pyloric stomach – pyloric caecae – Anus – feed primarily on sessile organisms Class Asteroidea Systems • Circulation – poorly developed with fluid filled chambers; – no heart; coelom ciliated for fluid movement • Excretion – no special organs – general diffusion across body surfaces like tube feet • Respiration – no special organs – across body membranes • Nervous System – associated with epidermis – circular oral nerve ring with branches into arms Asteroidea Body wall • • • • • Epidermis- outer surface; includes – mucous cells – epithelium – Pedicellariae- jawlike appendages of the epidermis • can open and close • used to clean body of debris or put debris on body Dermis- includes – nerve cells – connective tissue Skeleton- below dermis – made of ossicles – lattice like connections – Calcium carbonate – with spines and tubercles Muscle layer- below dermis Peritoneum that lines coelom Asteroidea Reproduction • are dioecious; external fertilization • usually 10 gonads; 2 in each arm • have fissiparitydivision of central disc into two animals Asteroidea Reproduction • free living larvae • bipinnaria- first larval form develops into • brachiolaria - shows development of arms Class Ophiuroidea Brittle Starfishes Class Ophiuroidea Brittle Starfishes and Basket Stars • 5 arms usually • central disc well marked off, no branches of gut in arms Class Ophiuroidea Brittle Starfishes and Basket Stars • no anus, no ambulacral groove • madreporite on oral surface • no suckers on tube feet, no ampullae (have a valve to control pressure) • no pedicellariae • able to move quickly and snake like hence their class name Class Echinoidea sea urchins, sea bisquits, sand dollars Class Echinoidea • no arms • skeleton is fused into a solid test • tube feet have suckers • covered with moveable spines and pedicellariae Class Echinoidea specialized mouth structures - Aristotle's Lantern Class Holothuroidea Sea cucumbers Class Holothuroidea Sea cucumbers • body elongated in oral-aboral axis • skeletal system reduced or absent • no spines or pedicellariae • mouth and anus at opposite ends of body Class Holothuroidea Sea cucumbers • no external madreporite • tube feet with suckers • respiration through anal respiratory tree • dioecious; single gonad • suspension or detritus feeders • commensal relationship with pearl fish Class Crinoidea Sea Lillies Class Crinoidea • most are extinct • most primative • all sessile, with stalk that attaches to substrate • have branched arms for filter feeding • no suckers on tube feet • no madreporite • no pedicellariae Sea Lillies