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Chapter 33 Phylum Porifera Sedentary suspension feeders, capturing food passed through the body Water into spongocel, out thru osculum Both vary in number Sequential hemaphrodites Eggs maintained in mesophyll, sperm out thru osculum Motile larvae find substrate to develop Celllular level of organization Choanocytes, or flagellated collar cells, line interior of spongocel to move water in Amoebocytes take food from water and choanocytes to digest it Produce antibiotic related compounds Classes of Porifera Class Calcarea Spicules, made by amoebocytes, of CaCO3 E.g. Grantia Class Hexacinellida Spicules, made of silica Referred to as ‘glass sponges’ Class Demospongiae Skeleton of silica spicules, flexible spongin, or both Referred to as ‘bath spongs’ E.g. Spongia Phylum Cnidaria Sessile and motile forms Diplobalstic and radially symmetrical Contain a gastrovascular cavity, from endoderm Cnidocytes capture prey Sac like body plan Contain nematocysts, Simplistic muscle and nerve which penetrate, stick tissues to, or tangle prey Gastroderm contracts with Polyp or medusa body closed mouth=shape change form coordinated by nerve net Some 1, other, or both in Arranged radially life Classes of Cnidarians Class Hydrozoa Class Scyphozoa Dominate medusa form and minor polyp stage All marine E.g. jellyfish Class Cubozoa Box-shaped medusa stage with complex eyes and potent venom Class Anthozoa Polyp stage only, mostly colonial and sessile All marine E.g. sea anemones and coral Class Hydrozoa Dominate polyp form and minor medusa stage Most marine, some freshwater Hydra Freshwater species only exhibit polyp form Can be motile Obelia Colony of chitinous covered polyps Portugese man-of-war Colony of polyps Original is air bladder and rest for feeding and reproduction Phylum Platyhelminthes Bilateral, tribloblastic, acoelomates with sac body plan Demonstrates cephalization and ladder like nervous system Organ system variations Hermaphroditic No specialized circulatory or respiratory organs Gastrovascular cavity branches throughout body to distribute material to cells Excretory system of flame cells for osmoregulation Reduced in parasitic forms Classes of Platyhelminthes Class Tubellaria Most marine, some freshwater, some terrestrial Specialized sense organs and nervous system Eyespot, auricle, pharynx, and varies reproductively E.g. planaria (Dugesia) Class Trematoda Parsitic, many with suckers to aid in attachment Body mostly reproductive organs Life cycle alters between sexual and asexual forms (intermediate host) E.g. Schistosoma, blood flukes, evade detection by changing surface proteins Class Cestoda Parasitic in vertebrates with scolex to attach to intestines Lack mouth and gastrovascular cavity, absorb through body Chains of proglottids, sacs of sex organs that fill with eggs and exit in feces Can form cyst stages to survive as larvae E.g. Taenia (dog/cat tapeworm) Phylum Nematoda Non-segmented, pseudocoelomates, covered by a cuticle Tube within a tube body plan characterized by an alimentary canal (mouth and anus) Lack a circulatory system Reproduce sexually with internal fertilization Male and female species separate and distinct in size Zygotes able to survive harsh conditions Longitudinal muscles = whip-like movement Nematode Examples Trichinella spiralis (trichinosis) Juvenile worms encyst in pig muscle, humans consume Adults burrow through intestines into lymph system Caenorhabditis elegans (C. elegans) Popular model for genetic research Enterobius vermicularis (pinworms) Common childhood disease Dinofilaria immitis (dog heartworms) Wucheria bancrofti (elephantiasis) Transmitted by mosquitos and live in lymph systems Necatur americanus (hookworms) Phylum Mollusca Mostly marine, some freshwater or terrestrial Soft-bodied, but secrete a shell of CaCO3 (some lost) Coelomates with 3 part body plan Foot: muscular organ for locomotion, attachment, or feeding Visceral mass: contains internal organs Mantle: suurounds visceral mass and may secrete shell; develops gills or lungs Feed via a radula Most separate sexes, but snails are hermaphrodites Open circulatory system, blood not confined to vessels Neural ganglia connected by nerve cord Classes of Molluscs Class Polyplacophora Body of dorsal plates, but unsegmented; no head but radula Class Gastropoda Herbivore’s uses radula to scrape, carnivorous to bore thru prey Developed head with eyes and demonstrates torsion Some hermaphroditic, but require another individual Class Bivalvia Two part shells secreted by mantle and controlled by muscles Can see growth rings; made of protein Little cephalization, no head or radula Gills for gas exchange, most suspension feeders Separate sexes Class Cephalopoda Active predators with beak like jaws and poisoned saliva Foot modified into a siphon to direct movement Closed circulatory system, well developed sense organs, and a brain Mantle covers visceral mass, but may be internal or missing Nautiluses are last surviving ammonites, posses shells Phylum Annelida Segmented worms, separated by partitions called septa Coelomates with a closed circulatory system Solid ventral nerve cord, anterior brain, and a ganglia in each segment Nephridia, coiled tubes for excretion in each segment Digestive stystem with pharynx, stomach, and intestines Some with setae, bristles, and parapodia, paddle-like appendages, for movement Classes of Annelids Class Polychaeta Each segment with parapodia and setae Marine animals that are mostly filter feeders, some predators Defined cephalization with eyes, sense organs, and jaws Class Oligochaeta Moist environments to allow gas exchange Include earthworms Hermaphroditic, posses a clitellum to aid cross-fertilization Class Hirudinea Most freshwater, some marine and terrestrial Invertebrate predators or parasites Slit skin with jaws or dissolve hole with enzymes Use anesthetic so undetectable and hirudin, an anticoagulant Phylum Arthropoda Specialized jointed appendages Hard exoskeleton of chitin and protein Sites for muscle attachment, protection, and stops desiccation Limits growth so must molt = energetically expensive Segmented bodies allows organ and system specializations Developed nervous system for sight, smell, and touch Open circulatory system Respiratory systems vary between tubes, lungs, and gills Subphyla of Arthropods Subphylum Crustacea Subphylum Chelicerata Mostly marine with gills Most collectively called Head with compound eyes & arachnids No antennae and simple eyes Specialized appendages 5 appendage pairs 2 pairs antennae 1 mandible and 2 mandible for feeding Walking legs on thorax Includes Isopods, Copepods and Krill, Barnacles, and Decapods 1 pair for feeding = chelicerae 1 pair sensory function = pedipalps 4 pair walking legs Book lungs for respiration Subphyla of Arthropods Subphylum Myriapoda Subphylum Hexapoda 1 Pair of antennae, 1 pair of Class Insecta 1 or 2 pairs of wings from thorax, cuticle extensions not appendages Advertize with colors , sounds, or odors for reproduction Metamorphosis to reduce madibles, and 2 pairs maxillae Class Diplopoda Millipedes are herbivores with 2 pairs of legs a segment Class Chilopoda Centipedes are carnivorous with 1 pair of legs a segment; poison claws on first segment competition within a species Complete (different and direct) or incomplete (similar and stages) Phylum Echinodermata Deuterostomes with a spiny endoskeleton covered by calcareous plates with spines Water vascular system includes tube feet for locomotion, feeding, and gas exchange Lack complex circulatory, respiratory, and excretory systems Internal and external parts radiate from center of organism Not true symmetry, larvae are bilateral and sieve plate offset in adult Lack cephalization, nervous system is ring with radial nerves in each arm Classes of Echinoderms Can regrow lost arms, turns stomach inside out to eat Long, flexible arms for movement Mouth is a jaw like structure, tube feet in rows Mouth faces up and arms for suspension feeding Lack spines, tube feet around mouth for feeding Armless with 5-sided organization, ringed by spines