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BEWARE ! ! BEWARE ! THE MOLLUSCS ARE COMING Phylum Mollusca • Very diverse group • There are 7 classes but we will only discuss 4 of them (Chitons (Polyplacophora), Cephalopods, Gastropods, Bivalves). • Some similarities between the groups can only be seen during development. • All mollusks have a free-swimming larval stage in their development – The Trochophore Larva Phylum Mollusca – Body Plan • Their body plans are very diverse. • This is why they are the second largest invertebrate phylum – what’s bigger? • Major features of the body plans in mollusks include: Foot (modified to tentacles in cephalopods) Mantle (tissue fold that covers the body) Shell (internal or external, CaCO3) Gills (outgrowths of the mantle wall) Visceral mass (guts) Radula (rasping tongue) Chitons – Eat algae or small animals from the surface of rocks. – Live near the shore or even in the deep ocean – Bottom surface is a muscular foot – Top surface is made of 8 overlapping plates of armor. Gastropods – Largest group of mollusks – Coiled shell – Some are soft bodied without a shell – The foot has a hard plate (operculum) on it that protects the body when it withdraws into the shell. – – – – – Clams, mussels, oysters, scallops. Two shells – “BIVALVIA” The halves are held together by strong muscles. Most can’t move – sedentary (except –SC) Filter feed with gills – food embeds in the mucus which is moved toward the mouth by cilia. – They have a stomach and waste leaves at the anus – near the excurrent siphon. – Mussels stick to rocks with a byssus (stringy, sticky threads) Cephalopods • • • • “head footed” Most complex Squid, cuttlefish, nautilus, octopi Nautilus have a complete external shell and many small tentacles that do not have suckers. • Octopi have 8 tentacles equipped with suckers, and they have completely lost their shell. • Squid have 10 appendages (8 arms + 2 tentacles with suckers on the tips – feeding). Also have an internal shell called a pen, that holds the shape of the body. NAUTILUS CUTTLEFISH OCTOPUS http://www.youtube.com/wa tch?v=5rqhomPaxhE This blue-ringed octopus has one of the most poisonous venoms in the world. Inhabit the Great Barrier Reef along the East Coast of Australia Cephalopods • • • • • Dioecious – males and females are separate. Internal fertilization Squid lay eggs and die Octopi will guard the eggs in a cave. To allow for quick locomotion, the cephalopods have incorporated a closed circulatory system. Cephalopods • Predatory – unlike the detritus-feeding or filterfeeding molluscs in the other classes. • Rapid locomotion requires an efficient respiratory system • The mantle contracts and pushes jets of water for the octopus and the squid to move. • Nautiloids swim with their tentacles. • Stomach and digestive system. • Closed circulatory system – vessels • 2 hearts – one for gill system and one for systemic. • Highly developed nervous system – brain – octopi can learn and remember. • Complex eyes – crucial to a predator. • http://www.youtube.com/watch?v=jsxAr_yrJtM Ecological Role • • • • • • Wide ecological distribution Aquatic and terrestrial Food source for many organisms Symbiosis Bivalves recycle sediment Shells provide homes for other animals when discarded. • Some are agricultural pests • Concentrate pollutants in their tissues (filter feed)