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Phylum Mollusca 27-4 Mollusks • • • • Phylum Mollusca One of the oldest and most diverse phyla Second largest phyla of animals Soft-bodied animals with an internal or external shell • Have a free swimming larval stage Body Plan • True coelomates • Complex, interrelated organ systems function together • 4 parts: foot, mantle, shell, visceral mass • Foot: used for crawling, burrowing, tentacles • Mantle: thin layer of tissue that covers most of the body • Shell: made by glands in mantle that secrete calcium carbonate (reduced or lost in some) • Visceral Mass- beneath mantle, consists of internal organs Feeding • Herbivores, carnivores, detritovores, parasites • Radula- flexible, tongue-shaped, rough structure – Used to scrape algae or eat soft tissues • Filter feeders- clams, oysters, scallops- siphon water – Siphon- tubelike structure for water to enter and exit Respiration, Circulation, Excretion • Gills in the mantle cavity- snails, clams and octopi • Through mantle blood vessels- land snails and slugs • Open circulatory system- snails, clam- slow because low oxygen demand • Closed circulatory system- octopi, squid- transports blood quickly • Cells release waste into blood and nephridia remove ammonia from the cell Response and Movement • Bivalves- burrow- simple nervous system – Chemical receptors, eye spots • Octopi- most highly developed nervous system in all invertebrates – Good memory, complex behavior • Snails move using mucus secreted along the base of the foot • Octopi and squid move using jet propulsion Reproduction • Sexually • External fertilization: sea snails and bivalves • Internal fertilization – Tentacled and some snails • Some are hermaphrodites - nudibranchs – Not self fertilizing Class Gastropoda • Pond snails, land slugs, sea hares, limpets and nudibranch • “Stomach-foot” • Single-shelled- protected by withdrawing and covering the opening with an operculum – Snails • No external shell- protected by hiding, ink squirting, poison, bright colors, nematocysts – Slugs, nudibranchs, and sea hares Nudibranch Class Bivalvia • Have two shells that are held together by one or two powerful muscles – Clams, mussels, oysters, scallops • Stay in one place, dig or burrow or attach • Scallops flap shells when threatened • Cilia or gills create current to circulate water • • • • • Class Cephalopoda Octopi, squid, cuttlefish, chambered nautilus “Head-foot” Most active of the mollusks Soft-bodied head attached to foot (tentacles) Tentacles have sucking disks and grab hold of prey • Most have an internal shell- except nautilus • Complex eyes- can distinguish shapes by sight and texture by touch Squid-color changes http://www.youtube.com/watch?v=PrnUIg302mw