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Chapter 28 Arthropods and Echinoderms Introduction to Arthropods • • • • • • “jointed feet” Most diverse and successful animals Over 750,000 species identified Segmented bodies Tough exoskeleton Jointed appendages Body Terms • Exoskeleton- external skeleton that protects and supports the body – Made of protein and a carbohydrate called chitin – Much variation on exoskeletons – Terrestrial arthropods (Ter. Arth.) have waxy covering to prevent water loss • Appendages- structures like legs and antennae that extend from the body wall Evolution has led to: • Fewer body segments – The body segments fused together • Highly specialized appendages – For feeding, movement, and more – Legs became: antennae, claws, wings, flippers, tails, mouthparts and walking legs Form and Function • Feeding • Varied eating habitsherbivores, carnivores, omnivores – Bloodsuckers, filter feeders, detritivores, parasites • Varied mouthpartspincers, fangs, jaws Respiration • Tracheal tubes- branching, air filled tubes that are in many ter. arth. • Spiracles- small openings alongside the body that allow air to enter and leave the tracheal tubes • Book lungs- organs with layers of respiratory tissue stacked ex) spiders • Gills- used by aquatic arthropods ex) crabs • Book gills- used by horseshoe crab Circulation • Open circulatory system- well developed heart pumps blood arteries tissues sinus collects around the heart and re-enters to be pumped through again Excretion • Malpighian tubules- saclike organs that extract wastes from blood and add them to feces to move through the gut – Used in ter. Arth. • Diffusion- moves cellular waste from the body to the water – Used in aquatic arth. Response • All have a brain • Well developed nervous system • Two nerves around the esophagus connect brain to the central nervous cord – Connects the ganglia which coordinate movement of legs and wings Movement • Well developed muscles controlled by nervous system • Individual muscles cells • Muscles generate force (to fly, walk, swim) by contracting a muscle and pulling on the exoskeleton Reproduction and Molting • Terrestrial – Internal fertilization • Aquatic – Internal and external fertilization • Molting – When an arthropod sheds its entire exoskeleton and makes a larger one in its place – Controlled by the endocrine system with hormones Subphyla of Arthropods • Crustacea – crabs, shrimp, crayfish • Chelicerata – spiders, ticks, scorpions • Uniramia – millipedes, centipedes • Class Insecta (no subphylum)– 3 part body, 3 pairs of legs 28-4 Echinoderms • “spiny skin” • Endoskeleton- hardened plates of calcium carbonate – Gives bumpy, irregular surface • Only live in the sea • No cephalization • Two sided- oral (with mouth)/aboral Echinoderm Characteristics • • • • • Spiny skin Internal skeleton Water vascular system Suction cup structure called tube feet 5 part radial symmetry – Larvae- bilaterally symmetrical • Deuterostomes Water Vascular System • Carries out essential body functions like respiration, circulation and movement • Madreporite- opening to the outside through which water passes – Connects to ring canal which extend out into radial canals along the 5 arms • Tube feet- suction cups on the underside of the body that help with movement and feeding Feeding • Urchins- scrape algae • Sea lilies- capture floating plankton • Sea cucumbers- take in sand and detritus off sea floor • Sea stars- use tube feet to pull open bivalves, push stomach out, pour enzymes and digest mollusks, brings back stomach into body Respiration and Circulation • Water vascular system • Thin walled tissues of tube feet for respiration • Skin gills gas exchange Excretion • Digestive wastes- feces through anus • Ammonia excreted through tube feet Response and Movement • Don’t have highly developed nervous system • Nerve ring around the mouth • Sensory organs that detect light • Tube feet • Endoskeleton structure/flexible joints Reproduction • External fertilization • Separate sexes Groups of Echinoderms • • • • • • • Urchins Sand dollars Brittle stars Sea cucumbers Sea stars Sea lilies Feather stars Ecology • Urchins control algae • Sea stars control clams and corals • Crown of Thorns – Feeds on corals – Rows of poisonous spines on arms – Great Barrier Reef damage