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Brain and Behaviour Second Hour – How do neurons communicate? Afferent (sensory) and Efferent (motor) nerves revisited •A demonstration •Interneurons Parts of the neuron Neuronal communication •Electrical and chemical Spatial and temporal summation Average Individual Reaction Time Measuring the speed of the nervous impulse .55 .50 .45 Shoulder with decision .40 .35 .30 .25 Ankle .20 .15 .10 Shoulder .05 1 2 3 Practice Trials Note: This is an approximation of your responses to show the pattern of response. Electrical communication within the neuron Myelin Sheath (speeds transmission) Axon Nodes of Ranvier Cell Body Dendrites (receivers) Axon Terminals (transmitters) Electrical Transmission Synapse Chemical transmission Electrical Transmission Neuron 1 Neuron 2 Electrical Transmission – The Action Potential At rest – Positive on the outside, negative on the inside 1. Action potential Sodium channels open - Positive sodium ions surge in making the inside of the neuron positive 2. Positive potassium ions are forced out. Excess positive sodium ions forced out. Stability returns 1. (outside) (inside) 2. Sodium+ Sodium+ Potassium+ Potassium+ (outside) (inside) (See Gleitman, p. 68) The action potential Voltage change over time Depolarization +Sodium in adds to +Potassium Inside voltage (mV) 40 resting potential Positive outside Negative inside Polarized Repolarization + Potassium out 20 0 -20 Hyperpolarization + Sodium out -40 -60 Resting + Potassium back in returns to positive outside negative inside -80 0 Stimulus onset 1 2 3 Time (milliseconds) refractory period 4 5 Electrical Transmission Synapse Chemical transmission Electrical Transmission Neuron 1 Neuron 2 (See Gleitman, p. 72) Axon of the sending neuron Neural impulse Axon terminal Presynaptic membrane inding site Postsynaptic membrane Synaptic vesicle Synapse (Synaptic gap or cleft) Neurotransmitter molecule Receiving neuron Chemical Transmission Electrical Transmission Synapse Chemical transmission Electrical Transmission Neuron 1 Summation Neuron 2 Temporal summation: several impulses from one source over time Dendrites Axon Hillock Point of summation Axon Spatial summation: several impulses from several sources converging at the same time