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Ecosystems The Rise of Life on Earth Ecosystems • Classification of Organisms – All organisms are classified base on their cell structure: • Eukaryotes • Prokaryotes Ecosystems • Classification of Organisms – Eukaryotic Cells • Cell membrane • Nucleus – Nuclear membrane – DNA Ecosystems • Classification of Organisms – Prokaryotic Cells • Bacteria • Cell membrane • No distinct nucleus Ecosystems • Classification of Organisms – Taxonomic Classification classification based on common characteristics Ecosystems • The Rise of Life on Earth – Requirements • Restricted temperature range • Liquid water Ecosystems • The Rise of Life on Earth – Reasons for these conditions • • • • • Earth’s distance from Sun Earth spins on tilted axis Earth’s size Internal heat engine Protective ozone layer Ecosystems • The Rise of Life on Earth – The rise of life on Earth is intimately linked to the physical and chemical evolution of the Earth Ecosystems • The Rise of Life on Earth – Two phases of development • Chemical evolution of organic molecules • Biological evolution of organisms Evolution of the Earth and Its Life forms Ecosystems • The Rise of Life on Earth – Chemical Evolution • Formation of oceans • Formation of primitive atmosphere • Formation of organic compounds necessary for life Ecosystems • The Rise of Life on Earth – Chemical Evolution • Formation of oceans – Volcanic “degassing” triggered start of hydrologic cycle Ecosystems • The Rise of Life on Earth – Chemical Evolution • Formation of primitive atmosphere – – – – – – CO2 N2 H 2O Methane (CH4) Hydrogen sulfide (H2S) Hydrogen chloride (HCl) Ecosystems • The Rise of Life on Earth – Chemical Evolution • Formation of organic compounds – Electrical discharges (lightning) – UV radiation – Geothermal energy Ecosystems • The Rise of Life on Earth – Chemical Evolution • Possible sites of formation of organic compounds – Atmosphere – Space – Mid-ocean hydrothermal vents Ecosystems • The Rise of Life on Earth – Chemical Evolution • Organic compounds probably collected in warm, shallow waters – Millions of years of chemical reactions gave rise to protocells capable of growing and dividing Ecosystems • The Rise of Life on Earth – Biological Evolution • Protocells evolved into prokaryotic anaearobic cells • Anaerobic cells underwent billions of years of evolution into a diversity of prokaryotic cells Ecosystems • The Rise of Life on Earth – Biological Evolution • No life on primitive Earth – No protective ozone layer to protect DNA – No evidence of fossils Ecosystems • The Rise of Life on Earth – Biological Evolution • The Oxygen Revolution – Photosynthetic algae in oceans – Evolution of ozone layer – Rise of life on land ~ 400 million years ago Ecosystems • Population Responses to Stress – Evolution – Adaptation – Natural Selection Ecosystems • Population Responses to Stress – Biological Evolution - a change in a population’s genetic makeup through successive generations – Theory of Biological Evolution - all lifeforms developed from earlier liofe-forms Ecosystems • Population Responses to Stress – Evidence of Earth’s life history • Fossils - mineralized of petrified remains of skeletons, bones, teeth, seeds, or impressions of these features • Paleontology - the study of fossilized organisms Ecosystems • Population Responses to Stress – Evidence of Earth’s life history • Fossil record is incomplete – Fossil record represents only 1% of species believed to have existed in the past Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Gene - segments of DNA molecules found in chromosomes; coded units of information about specific traits that are passed from parents to offspring during reproduction Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Gene Pool - the sum total of all genes found in individuals of a population of a particular species Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Members of a population generally have the same number and kinds of genes • Allelle - slightly different molecular forms found in a particular gene Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Different combinations of alleles are inherited so that different members of a population have genetic diversity Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Mutation - random change in the structure or number of DNA molecules in a cell Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Causes of mutations – – – – Radioactivity X-rays Chemicals (mutagens) Random mistakes during DNA replication Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Types of mutations – Harmful – Beneficial Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Beneficial Mutation - a mutation that results in new genetic traits that give their bearer and its offspring better chances for survival – Adaptive Trait - a genetically-controlled trait that helps an organism survive and reproduce under a given set of environmental conditions • Development of coloration that allows prey to hide from predators Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Differential Reproduction - a phenomenon in which individuals with adaptive genetic traits produce more living offspring than do inviduals without such traits Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Natural Selection - the process by which a particular beneficial gene (or set of genes) is reproduced more than other genes in succeeding populations. Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • The end result of natural selection is a population that contains a greater proportion of organisms better adapted to certain environmental conditions Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Limits to Adaptation – A change in environmental conditions can only lead to the adaptation for traits already present in the gene pool Ecosystems • Population Responses to Stress – The Theory of Biological Evolution • Limits to Adaptation – Even if an adaptive trait is present in a population, the population’s ability to actually adapt to a change in environmental conditions can be limited by its reproductive capacity • Reproductive lag time • Example: r-strategists vs K-strategists Ecosystems • Population Responses to Stress – Evolution of Species (Speciation) • Two components of speciations – Geographic Isolation – Reproductive Isolation Ecosystems • Population Responses to Stress – Evolution of Species (Speciation) • Two components of speciation – Geographic Isolation - separation of populations of a species for a long time into areas with different environmental conditions • Separation due to physical barriers • Separation due to natural disasters • Separation by wind and oceanic currents Ecosystems • Population Responses to Stress – Evolution of Species (Speciation) • Two components of speciation – Reproductive Isolation - long-term geographic separation of members of a particular sexually reproducing species Ecosystems • Population Responses to Stress – Evolution of Species (Speciation) • Two components of speciation – Reproductive Isolation • Occurs when mutations and natural selection operate independently between to geographically isolated populations • Results in divergence (divergent evolution) and speciation Ecosystems • Population Responses to Stress – Extinctions • Causes – Continental drift – Climate change • Gradual • Catastrophic Ecosystems • Population Responses to Stress – Extinctions • Types – Background Extinctions • Caused by changes in local conditions – Mass Extinctions • Abrupt rise in extinction rate above background levels Ecosystems • Population Responses to Stress – Extinctions • Mass extinctions are followed by periods of recovery called adaptive radiations in which new species fill new or vacated niches • Extinction of dinosaurs allowed for the radiation of mammals Ecosystems • Community and Ecological Responses to Stress – Ecological Succession - the process in which communities of plant and animal species in a particular area are replaced over time by a series of different and often more complex communities Ecosystems • Community and Ecological Responses to Stress – Types of succession • Primary Succession - sequential development of communities in a bare area that has never been occupied by a community of organisms – Pioneer Species - first plants to colonize a lifeless area; typically r-strategists Ecosystems • Community and Ecological Responses to Stress – Types of succession • Secondary Succession - sequential development of communities in an area in which the natural vegetation has been removed or destroyed but the soil is not destroyed Ecosystems • Human Impacts on Ecosystems – Simplification of natural ecosystems – Enhancing populations of pest species – Elimination of predators – Introduction of exotic species – Overuse of potentially renewable resources – Interference with biogeochemical cycles