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Chapter 37 Plant Nutrition PowerPoint Lectures for Biology, Seventh Edition Neil Campbell and Jane Reece Lectures by Chris Romero Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Overview: A Nutritional Network • Every organism continually exchanges energy and materials with its environment • For a typical plant, water and minerals come from the soil, while carbon dioxide comes from the air Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Concept 37.1: Plants require certain chemical elements to complete their life cycle H2O CO2 • Plants derive most of their organic mass from the CO2 of air, but they also depend on soil nutrients such as water and minerals O2 O2 Minerals CO2 H2O Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Macronutrients and Micronutrients • A chemical element is considered essential if it is required for a plant to complete its life cycle • Researchers use hydroponic culture to determine which chemicals elements are essential Control: Solution containing all minerals Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Experimental: Solution without potassium • Nine of the essential elements are called macronutrients because plants require them in relatively large amounts • The remaining are called micronutrients because plants need them in very small amounts Symptoms of Mineral Deficiency • Symptoms of mineral deficiency depend on the nutrient’s function and mobility within the plant • The most common deficiencies are those of nitrogen, potassium, and phosphorus Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Healthy Phosphate-deficient Potassium-deficient Nitrogen-deficient Texture and Composition of Soils • Topsoil is a mixture of particles of rock, living organisms, and humus (the remains of partially decayed organic material) A B C Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Animation: How Plants Obtain Minerals from Soil • After a heavy rainfall, water drains from the larger spaces of soil, but smaller spaces retain water because of its attraction to clay and other particles Soil particle surrounded by film of water Soil particle Root hair Water available to plant Root hair Air space Soil water Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cation exchange in soil Concept 37.4: Plant nutritional adaptations often involve relationships with other organisms • Two types of relationships plants have with other organisms are mutualistic: – Symbiotic nitrogen fixation, involving roots and bacteria – Mycorrhizae, involving roots and fungi Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings The Role of Bacteria in Symbiotic Nitrogen Fixation • Symbiotic relationships with nitrogen-fixing bacteria provide some plant species with a built-in source of fixed nitrogen • For agriculture, the key symbioses between plants and nitrogen-fixing bacteria occur in the legume family (peas, beans, and other similar plants) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Along a legume’s roots are swellings called nodules, composed of plant cells “infected” by nitrogen-fixing Rhizobium bacteria 5 µm Bacteroids within vesicle Nodules Roots Pea plant root. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Bacteroids in a soybean LE 37-11 Infection thread Rhizobium bacteria Dividing cells in root cortex Bacteroid Infected root hair Dividing cells in pericycle Developing root nodule Bacteroid Bacteroid Nodule vascular tissue Symbiotic Nitrogen Fixation and Agriculture • Crop rotation takes advantage of the agricultural benefits of symbiotic nitrogen fixation • A non-legume such as maize is planted one year, and the next year a legume is planted to restore the concentration of nitrogen in the soil Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Mycorrhizae and Plant Nutrition • Mycorrhizae are mutualistic associations of fungi and roots • The fungus benefits from a steady supply of sugar from the host plant • The host plant benefits because the fungus increases the surface area for water uptake and mineral absorption Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings The Two Main Types of Mycorrhizae • In ectomycorrhizae, the mycelium of the fungus forms a dense sheath over the surface of the root Epidermis Cortex Mantle (fungal sheath) 100 µm Endodermis Fungal hyphae between cortical cells Mantle (fungal sheath) Ectomycorrhizae. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (colorized SEM) • In endomycorrhizae, microscopic fungal hyphae extend into the root 10 µm Cortex Epidermis Cortical cells Endodermis Fungal hyphae Vesicle Casparian strip Root hair Arbuscules (LM, stained specimen) Endomycorrhizae. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Epiphytes, Parasitic Plants, and Carnivorous Plants • Some plants have nutritional adaptations that use other organisms in nonmutualistic ways Video: Sun Dew Trapping Prey Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings LE 37-13a Staghorn fern, and epiphyte. This tropical fern (genus Platycerium) grows on large rocks, cliffs, and trees. It has two types of fronds: branched fronds resembling antlers and circular fronds that form a collar around the base of the fern. LE 37-13b Host’s phloem Dodder Haustoria Mistletoe, a photosynthetic parasite. Dodder, a nonphotosynthetic parasite. Indian pipe, a nonphotosynthetic parasite. LE 37-13c Venus’ flytrap. Pitcher plants. Sundews.