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THE TREE OF LIFE WHY CLASSIFY? • We classify to make organisms easier to study. • To study the diversity of life, biologists use a classification system to name organisms and group them in a logical manner. • The field of taxonomy allows scientists to classify organisms and assign each organism a universally accepted name in order to study the diversity of life. LINNAEUS’S SYSTEM OF CLASSIFICATION • During the eighteenth century, Swedish botanist Carolus Linnaeus developed a two-word naming system called binomial nomenclature • Linnaeus often classified organisms based on similarities in structures and details of anatomy BINOMIAL NOMENCLATURE • In binomial nomenclature, each species is assigned a 2-part Latin name • The name is always written in italics with the 1st word is always capitilized • Example: Homo sapiens (humans) • The 1st part of the name is the genus and the second part of the name is unique to each species within that genus LINNAEUS’S SYSTEM OF CLASSIFICATION • Linnaeus’s system of classification uses 7 taxonomic categories (from largest to smallest): • Kingdom – largest & most inclusive • Phylum – includes many different organisms that share important characteristics • Class – composed of similar orders • Order – composed of similar families • Family – genera that share many characteristics • Genus – a group of closely related species • Species – a group of individuals so similar that they can breed and produce fit offspring HUMAN CLASSIFICATION • Kingdom: Anamalia • Phylum: Chordata • Subphylum: Vertebrata • • • • • Class: Mammalia Order: Primates Family: Hominidae Genus: Homo Species: Homo sapiens KINGDOM Animalia PHYLUM Chordata CLASS Mammalia ORDER Carnivora FAMILY Ursidae GENUS Ursus SPECIES Ursus arctos PROBLEMS WITH TRADITIONAL CLASSIFICATION • During Linnaeus’s time, scientists classified organisms based on their physical appearance…but today…we know that doesn’t always work. • Ex: Dolphins Fish or Mammals? • Remember Convergent Evolution sometimes organisms that are different from each other evolve similar body structures, due to the change in the environment. • This does not mean they need to be classified in the same group…maybe they only evolved similar body structures! • These situations make it very difficult for scientists to classify. EVOLUTIONARY CLASSIFICATION • Biologists now group organisms into categories that represent lines of evolutionary descent, not just physical features • Evolutionary classification (phylogeny) is the strategy of grouping organisms together based on their evolutionary history TRADITIONAL V/S EVOLUTIONARY CLASSIFICATION CLASSIFICATION USING CLADOGRAMS • To refine evolutionary classification, biologists now prefer a method called cladistics • Cladistics considers only those characteristics that are new characteristics that arise as lineages evolve over time • Characteristics that appear in recent parts of a lineage but not in its older members are called derived characters • Characteristics that are shared by all members of a lineage are ancestral/primitive characters and should not be used when building cladograms. • Cladogram – a diagram that shows the evolutionary relationships among a group of organisms; includes new characteristics that arise as lineages evolve. DERIVED CHARACTERISTICS ANATOMY OF THE CLADOGRAM Monophyletic group = clade J Taxon I F G H C D E Paraphyletic group Lineage (represents a sequence of ancestor-descendent populations) A B Node (represents most recent common ancestor of two or more taxa) Synapomorphy (shared derived character) MONOPHYLETICGROUPS: include ancestor & all descendents PARAPHYLETIC GROUPS: includes ancestor and some, but not all descendent MONOPHYLETIC CLADES • A valid clade is monophyletic signifying that it consists of the ancestor species and all its descendants. OUTGROUPS • Systematists use a method called outgroup comparison • To differentiate between shared derived and shared primitive characteristics • As a basis of comparison we need to designate an outgroup • which is a species or group of species that is closely related to the ingroup, the various species we are studying • Outgroup comparison • Is based on the assumption that homologies present in both the outgroup and ingroup must be primitive characters that predate the divergence of both groups from a common ancestor PERFORMING OUTGROUP COMPARISON OUTGROUP COMPARISONS • First, look at the animals we are studying and establish which characteristics that they share & which are unique to each individual species. • Then use these derived characters to build your cladogram. CELLS Slug Catfish Frog Tiger Human BACKBONE LEGS HAIR OPPOSABLE THUMB THE THREE-DOMAIN SYSTEM • Molecular analyses have given rise to the most current classification system – the Three Domain System • The 3 Domain System is the most recent classification system and includes: • Bacteria • Archaea • Eukarya DOMAINS ARE BIGGER/MORE INCLUSIVE THAN KINGDOMS Domains Kingdoms Bacteria Eubacteria Archaea Archaebacteria Eukarya Protista Fungi Plantae Animalia These 2 kingdoms used to be combined into one called “Monera” REMEMBER - THERE ARE 3 METHODS OF CLASSIFYING ORGANISMS • The 3 Domain System (most current) • Bacteria, Archae, & Eukarya • The 5 Kingdom System (Whittaker) • Monera, Protista, Fungi, Plantae, Animalia • The 6 Kingdom System – still used in conjunction with the 3 domain system • Archaebacteria, Eubacteria, Protista, Fungi, Plantae, Animalia Section 18-3 Classification of Living Things Go to Section: DOMAIN BACTERIA • • • • • • Kingdom: Eubacteria Unicellular Prokaryotic Cell wall of peptidoglycan Autotrophic or heterotrophic Examples: Streptococcus, E. Coli DOMAIN ARCHAEA • • • • • • Kingdom: Archaebacteria Unicellular Prokaryotic Cell walls w/out peptidoglycan Autotrophic or heterotrophic Methanogens & halophiles DOMAIN EUKARYA WHITTAKER’S FIVE-KINGDOM SYSTEM OUR CHANGING VIEW OF BIOLOGICAL DIVERSITY THE THREE DOMAIN SYSTEM Describes classification as: • • • • • • • Not all prokaryotes are closely related (not monophyletic) Prokaryotes split early in the history of living things (not all in one lineage) Archaea are more closely related to Eukarya than to Bacteria Eukarya are not directly related to Eubacteria There was a common ancestor for all extant organisms (monophyletic) Eukaryotes are more closely related to each other (than prokaryotes are to each other)