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UV and Insect eyes LIGHT & PHOTOSYNTHESIS Spectrum Fate of intercepted light? Reflect Transmit Absorb Fate of leaf intercepted light? Reflect - (6-12% PAR, 70% infrared, 3% UV) Degree of reflection varies with type of leaf surface Transmit - average 10-20% (primarily green and far red) What would influence this? thickness and structure of leaf Absorb – What does this depend upon? Fig. 5.4 Absorption What determines how much light a plant or a group of plants will intercept? Quantity and position of leaves Leaf area index Leaf Area Index LAI Interpretation of LAI see fig 5.5 (d) pg 47 (m2 leaf area/m2 ground area) LAI 3 means 3m2 leaf area over each m2 of ground LAIs of different canopy types Summer Temperate Deciduous Forest: LAI 3-5 (1-5% light hitting canopy reaches floor) Summer Pine Forest LAI 2-4 (10-15% light hitting canopy reaches floor) Tropical Rain Forest LAI 6-10 (.25 – 2% light reaches floor) Light levels and LAI Why is the amount of light reaching the forest floor of the pine forest greater than that of the deciduous forest? Does the LAI of an area change throughout the year? What are the consequences of this for plants of the forest floor? Light Levels Are there other factors other than LAI and leaf angle that would influence the amount of light a forest floor organism might receive? See Fig. 5.7 page 49 Light levels: year profile Photosynthetic Terms PAR – photosynthetically active radiation Light compensation point (LCP) Light saturation point (LSP) photoinhibition PS terms expanded Light compensation point: rate of C02 uptake in PS = rate of C02 loss in Respiration Short version rate of PS = rate of respiration Light Saturation Point – Point at which increasing light does not increase PS Photoinhibition - High light levels inhibit PS LCP Shade –tolerant Low PS rates Low respiration rates Low metabolic rates Low growth rates Shade -intolerant Higher PS rates Higher respiratory rates Higher growth rates Lower survival in shade conditions Shade and Seedlings Apply your Smarts How does the concentration of Rubisco relate to photosynthesis rate? What does the production of Rubisco have to do with respiration? What do respiration rates have to do with tolerance to shade? Apply your Smarts What do respiration rates have to do with the light compensation point? What does the light compensation point have to do with shade tolerance or intolerance? In general what types of plants would you expect to have a lower light saturation point: shade inotlerant or shade tolerant Why? Apply your Smarts What would you change in a plant that would increase or lower the light compensation point? Plant a shade intolerant plant in the shade. What would the plant do to compensate for being in low light? Leafs and Light How does the amount of light a leaf receives influence its shape and sizE? See page 52 And what does surface area and volume have to do with it? Shade-Sun leaf PS and Temperature See page 59 Fig. 6.3-6.4 PS and Temperature PS and temp Plants and Temperature Heat gain and loss – What are the parameters? Reflectivity of leaf and bark Orientation of leaves to sun and wind? Size and shape of leaves Frost hardiness – Transpiration cooooooling……… Plants and Temperature Frost hardiness – mostly genetic Addition of protective compounds (antifreeze type) allows super cooling Ice forms in the cell wall???? Transpiration cooooooling……… Temperature and plants Photosynthesis Light Absorption PS Activity Problem 100% Relative PS activity 0% 400 500 600 Wavelength 700 Light Problem A B C D 100% Relative Absorbency 0% 400 500 600 700 Wavelength (nm) Photosynthesis 6CO2 + 12 H20 ---> C6H12O6 + 6H20 + 602 OR 6CO2 + 6 H20 ---> C6H12O6 + 602 PS Problem If you put a plant in a closed container with an atmosphere of 14C02, and H2180 where would you find the radioactive carbon and the heavy oxygen (180) in the container and/or plant? Fates of reactants H 20 C6H1206 02 H20 H 20 C02 Movement of water