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Systems Approach Ecosystem Community Population Landscape Ecosystem Community -Populations of different species that interact. Ecosystem - the various communities of organisms interacting with the environment. Landscape -a physical area such as a watershed consisting of various landscape elements (fields, forests) and the species found. Community • • • • • • Species richness Species diversity Functional groups Life Forms Trend Analysis Statistical Approach Ecosystem • Energy Flow • Nutrient cycling Landscape • Spatial Arrangement of landscape elements or units. Community • Central Question: How is a community structured and by what? • Are there patterns in community structure? • If so, what has caused this pattern • • • • Competition Predation Time Size of patches (islands) Equilibrium Non-equilibrium Equilibrium - focus on the mean Non-equilibrium - focus on the variation Equilibrium Non-equilibrium Mean Predictability Variation Unpredictable Competition Island Biogeography Succession Disturbance Competition • An equilibrium concept. Data from Gause 1934 • Competition is often not an important structuring process in communities. Species area relationship • Preston (1948, 1960 and 1962) – S = CAz – log S = (log C) + z(log A) Island Biogeography • 2 processes - extinction and colonization • Function of – island size and distance from mainland. Succession Disturbance a non-equilibrium concept -fire, insect damage, wind, flooding -unpredictable in space and time -results in non-equilibrium dynamics -recognized as being central to ecosystems Modeling Disturbance Intermediate Disturbance Hypothesis Predator Mediated Co-existence Hyp. Effect of grazing on species richness Succession In the Boreal Forest Holling’s Four-Box Model of ecosystem function. Keystone Species Other Keystone species: - Beaver (create ponds) - Elephants (Trees) - Sea otter (Seaurchins) Community structure • Equilibrium and non-equilibrium AND Time. Complexity Stability Stability Complexity Complexity Stability Debate • Elton Complexity -> Stability • May 1970’s Modeling – Not so! • Subsequently refuted (late 80’s & 90’s) • Yodzis - modeling – Tillman – experiments – BUT we still did not understand the nature of stability • Populations Summary – movement – Fragmentation – Spatially structured populations • Communities – Dynamic equilibrium/non-equilibrium – Competition – Island Biogeography • Protected areas - corridors – Disturbance • Intermediate disturbance hypothesis • predator mediated co-existance • Ecosystem stability