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Population Growth (Ch. 11) Population Growth • 1) Geometric growth • 2) Exponential growth • 3) Logistic growth Geometric Growth • Growth modeled geometrically – Resources not limiting – Generations do not overlap • Recall: • 1) = Nt+1 / Nt • 2) = Ro Geometric Growth • Growth modeled geometrically – Resources not limiting – Generations do not overlap • Equation: Nt = No t – – – – Nt = Number inds. @ time t No = Initial no. inds. = Geometric rate of increase t = Number time intervals Geometric Growth • Phlox (annual plant) Fig. 11.3 Exponential Growth • Growth modeled exponentially – Resources not limiting – Generations overlap • Recall: 1) Per Capita Rate of Increase (r) 2) r = (ln Ro) / T 3) r = b – d Exponential Growth • Equation: dN / dt = rmax N • dN / dt means “change in N per unit time” • Recall r: per capita rate of increase (Ch. 10) • rmax: Special case of r (intrinsic rate of increase). b - d under optimum conditions – b = birth rate and d = death rate (rates per individual per unit time) • As N increases, dN/dt gets larger. Exponential Growth • For exponential growth: Nt = N0 ermaxt • • • • • Nt = No. inds. at time t. N0 = Initial no. inds. e = Base natural logarithms rmax = Intrinsic rate of increase t = Number time intervals Resources not limiting Geo: Nt = No t Exponential Growth: Example • Whooping crane Exponential Growth: Example • Hunting/habitat destruction • Federally listed Endangered(1967). Down to 22!! Fig. 11.6 Exponential Growth: Example • Raising birds: costumes avoid imprinting on humans Exponential Growth: Example • Teaching young birds to migrate (Wisconsin to Florida) 1996 Geometric or Exponential? Nt = N0 ermaxt Nt = No t Geometric Exponential Generations Geometric or Exponential? Nt = No t Nt = N0 ermaxt Geometric Exponential Separate Generations Overlap λ=1 Population stable r=0 λ>1 Population growing r>0 λ<1 Population shrinking r<0 Logistic Population Growth • Unlimited resources?? nothing lasts forever… • As resources depleted: logistic population growth. Logistic Population Growth • As resources depleted, logistic population growth (generations overlap) – Sigmoid (S-shaped) curve. – Carrying capacity (K): number environment can support. Fig. 11.8 Ex Logistic Population Growth • Yeast growth (limited alcohol) • Max. 17% (34 proof) Fig. 11.9 Logistic Population Growth Equation: dN/dt = rmax N (1-N/K) • rmax = Intrinsic rate of increase (ideal conditions) • N = population size @ time t • K = carrying capacity • Or: dN/dt = rmax N (K-N) K How does this work? • dN/dt = rmax N (K-N) K • N small: rmax N (K-0) or ≈ rmax N (1) K At small N, acts like exponential growth! • N big: rmax N (K-K) or ≈ rmax N (0) K At larger N, growth slows: stops at K Logistic Population Growth 1-N/K is “scaling factor” When N nears K, dN/dt nears zero. Fig. 11.13 Logistic Population Growth • r: actual (realized) reproductive rate (b-d) • Max. @ small N • When N=K, r=0 • So b=d and b-d=0 • Above K? r negative Fig. 11.14 dN/dt = rmaxN (1-N/K) Concepts! • Population growth (# added per unit time) highest when N=K/2 • Maximum sustainable yield: largest sustainable harvest Concepts! • N/K: reflects environmental resistance – Factors that limit population size Environmental resistance – Density-dependent factors: depend on density (N/K) • Disease, Resource competition – Density-independent factors: not related density • Natural disasters (hurricane, fire, flood) dN/dt = rmaxN (1-N/K) Organism Size and Population Density • A search for patterns…… Size – Size vs. density (neg. correlation) – Generation time vs. size? Gen time Generation time vs. size rmax • Positive correlation • Log-log scale size vs. rmax? Size rmax vs. size • Negative correlation • Note log:log scale Human Population • How many? • Where? • Age distributions and growth potential How many? • • • • • • Human Population How many? 7.09 billion (6/17/13) 7.02 billion (6/11/12) 6.925 billion (6/19/11) 6.448 billion (6/18/05) Check it out now at: • http://www.census.gov/ipc/www/idb/worldpopinfo.php Where (continent)? • Where? Human Population Fig. 11.23 Fig. 11.22 Human Population • Where? Human Population • Age distributions and growth potential, 2008 Fig. 11.24 Human Population • “Population bomb”: potential of population to explode as people age 2000/2001-Present - New Silent Generation or Generation Z 1980-2000 - Millennials or Generation Y 1965-1979 Generation X 1946-1964 - Baby Boom 1925-1945 – Silent Generation 1900-1924 – G.I. Generation Human Population • Human pop. curve: shape? • What Earth’s K for humans? Human Population • Depends in part on lifestyle! – Ecological footprint: resource use – Biocapacity: resource supply – Deficit if use>supply: US largest deficit Course sequence • • • • • • Natural history (Done!) Individuals (Done!) Populations (Done! Except Life Histories) Species interactions (You are here!) Communities/ecosystems Geographic/global ecology 5 main types of interactions among species: Effect on species A Effect on species B Competition - - Predation + - Parasitism + - Commensalism + 0 Mutualism + + Type of interaction Species Interactions: Competition (Ch. 13) Competition (Ch. 13) • Definition: – Individuals attempt to gain more resource in limiting supply – (-,-) interaction: both participants get less • Intraspecific: Within species. • Interspecific: Between species. Competition • Interference Competition: – Individuals interact with each other • Resource (Exploitation) Competition: – Individuals interact with resource