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Chapter 56 Community Ecology Why knowing about ecosystems is important? Scientific knowledge as ‘models’ Basic ecological concepts Biotic interactions in an ecosystem Question Chapter 56 -- Community Ecology 1 Why understanding ecosystems is important DDT (dichloro-diphenyl-trichloroethane) 1874: discovery 1934: Paul Müller, Swiss chemist “polychlorinated hydrocarbon” DDT is a potent insecticide ... Low toxicity for humans (mammals) Many Agricultural uses e.g., corn earworm, cotton bollworm tobacco budworms Control of diseases malaria, typhus WWII Dutch elm disease Cotton bollworm Chapter 56 -- Community Ecology 2 By the 1950s other people were seeing other, unintended effects . . . Fish -- trout, sunfish, suckers, etc -- higher toxicity than to mammals Birds of prey -- eagles, osprey, hawks, etc -- Reproductive effects Songbirds What was impact of Rachel Carson’s “Silent Spring”? -- 1962 Tranforms U.S. environmental consciousness pairs (in 1000s) Nat. Envr. Prot. Act (NEPA) CAA & CWA 7 DDT banned in 1973 6 5 4 Number of bald eagle pairs in lower 48 states 3 2 1 0 1960 Question Chapter 56 -- Community Ecology 1970 1980 1990 2000 3 Why did we misjudge effects of DDT? An incomplete understanding of ecosystems . . . • • • • How do organisms interact with their surroundings? How do organisms interact with each other? How do nutrients (and pollutants) move & change in an ecosystem? How do ecosystems change over time? Our answers to these form a ‘model’ of ecosystem function What is a scientific ‘model’? How does it differ from traditional ‘fact-based’ knowledge? 1) Recognized to be incomplete 2) simplified version of reality 3) explain all current information 4) are modified over time Chapter 56 -- Community Ecology 4 A ‘food chain’ and ‘food web’ illustrate two models with different levels of detail Chapter 56 -- Community Ecology 5 Some Basic Concepts What is an Ecosystem? “A community of organisms and the physical surroundings with which they interact.” What are the two components of an ecosystem? -- “Biotic” vs “abiotic” What are some abiotic factors that define a habitat? What are 3 levels at which biologists study the organisms of an ecosystem? Species: Population: Community: Question Chapter 56 -- Community Ecology 6 How big is an ecosystem? Depends on model being studied . . . microecosystems organisms forests biomes Why are ecosystems said to be “open systems”? Chapter 56 -- Community Ecology 7 What is an ecological niche? “ The sum of all the resources an organism requires” What is a ‘fundamental’ niche? … ‘realized’ niche? How do species interact within an ecosystem? Competition Predator-prey Symbiosis Chapter 56 -- Community Ecology 8 Competition What are two forms of competition? Intraspecific competition Interspecific competition Interactions cascade through an ecosystem Why did removing seed-eating rodents from desert ecosystem favor then diminish ant populations? Question Chapter 56 -- Community Ecology 9 What happens when two species compete for the same resources? What is the “Competitive Exclusion Principle”? Classic Paramecium experiment How does “Resource Partitioning” prevent competitive exclusion? e.g. Caribbean anoles Question Chapter 56 -- Community Ecology 10 Why does introduction of exotic species often disrupt ecosystems? Examples: Zebra mussels Japanese knotweed Grey Squirrel Emerald Ash Borer Chapter 56 -- Community Ecology 11 Predator-Prey relationships Many animal defense mechanisms Vigilance, e.g., Flocking ostriches Concealment, e.g., Chuckwalla lizard Warning Coloration, e.g., Skunk Camouflage (cryptic coloration) Chapter 56 -- Community Ecology 12 Animal defense mechanisms, con’t Mimicry Batesian mimicry Müllerian mimicry How are mimicry and cryptic coloration different? Chapter 56 -- Community Ecology 13 Some other really neat defense mechanisms. . . Read National Geographic article on reserve in library “The Art of Deception” By Natalie Angier http://mclib.marietta.edu/search~S0?/rbiol+101/rbiol+101 /1,1,1,B/l856~b1233151&FF=rbiol+101&1,1,,1,0/startrefe rer//search~S0/rbiol+101/rbiol+101/1,1,1,B/frameset~1233 151&FF=rbiol+101&1,1,/endreferer/ Self-quiz is available Chapter 56 -- Community Ecology 14 Predators evolve strategies also Predators and prey “co-evolve’ Angler fish Snake Common stargazer fish Chapter 56 -- Community Ecology 15 What are the types of symbiosis? Mutualism Commensalism Parasitism external internal These relationships are not always what they seem… Chapter 56 -- Community Ecology 16 What type of symbiosis does each of these relationships represent? How is each organism benefited or harmed? Rhino and tick bird Mistletoe on pine Chapter 56 -- Community Ecology 17 Let’s test our knowledge… Symbiosis Question Defense mechanisms Question 1 Question 2 Types of Mimicry Question Chapter 56 -- Community Ecology 18 Species Distribution (from chapter 55.2) You are not responsible for: -- logistic growth curve equation -- r & k selected species Geographic range How are populations dispersed? What factors determine how individuals are dispersed? “Clumped” Chapter 56 -- Community Ecology 19 What causes population growth cycles? (from chapter 55.2) e.g., central European larch bud moth -- periodic outbreaks every 8-10 years Similar cycles occur here for tent caterpillars Question Zimmer, 1999. Science 284:86; Ranta, et al, 2002. Science 298:973 Chapter 56 -- Community Ecology 20 Ecosystems change geographically Gradients of physical conditions Create an “Ecotone” Chapter 56 -- Community Ecology 21 Read about human population growth (Chapter 55.7) Exponential trends Population pyramids Doubling times Chapter 56 -- Community Ecology 22 How do Ecosystems change over time? Succession primary vs secondary pioneer species climax community Why does succession occur? Chapter 56 -- Community Ecology 23 Secondary Succession Disturbances -- Fire in Yellowstone Chapter 56 -- Community Ecology 24 Old Field Succession Question Chapter 56 -- Community Ecology 25